3065. Zoology is Zoogeny divided and self-substantially
represented. What in Zoogeny was the organ of
a single indivisible animal here becomes the organ of a
separated animal, or becomes a self-substantial animal.
3066. The self-substantial animals are only parts of
the Great animal, which is the Animal Kingdom.
3067. The animal kingdom is only one animal, i. e.
the representation of animality with all its organs, each
of which is a whole for itself.
3068. A single animal originates, if a single organ
frees itself from the general animal body, and yet exercises
the essential animal functions.
3069. The animal kingdom is only a dismemberment
of the highest animal, i. e. of Man.
3070. Animals become nobler in rank, the greater the
number of organs which are collectively liberated or
severed from the Grand animal, and which enter into combination.
An animal, which e. g. lived only as intestine,
would be, doubtless, inferior to one which with the intestine
were to combine a skin; and that animal again
must be regarded as higher than the latter, which should
present, in addition to these organs, vessels, liver,
branchiæ, tracheæ, and lastly bones, &c.
3071. Animals are gradually perfected, entirely like
the single animal body, by adding organ unto organ.
The animal kingdom is developed through the multiplication
of the organs.
3072. Each animal ranks therefore above the other;
two of them never stand upon an equal plane or level.
Animals are distinguished by their position of stages or
degrees from each other, by the number of their different
organs, but not by the division of a single organ.
3073. The animal system cannot be arbitrarily disposed
according to this or that organ, just as it may
chance to meet the eye; but only in accordance with the
rigid prescripts of the animal body's genesis.
3074. The animal body separates into two series of
organs, which, corresponding with, pursue a proximal
course in relation to, each other; into the Anatomical
systems and the Sensorial organs, unto which the sexual
parts appertain.
3075. The number of the sensorial organs is 5, and
they thus stand according to their genetic development
one above the other:
Tactile sense or Skin.
Gustatory sense or Tongue.
Olfactory sense or Nose.
Auditory sense or Ear.
Optical sense or Eye.
3076. In animals, which are characterized by the
sense of feeling or touch, the other sensorial organs must
be either still wanting, or if present but imperfectly conditioned,
i. e. not constituted like those of man, who is
the type, pattern, or paragon for every formation.
3077. Their sensations are limited to those of general
touch or feeling, and of those derived through the medium
of the other senses we meet with but feeble manifestations.
3078. Their body itself will only be a tegumentary
body, with the organs subordinated to the integument,
namely, the viscera. They are therefore devoid of a true
tongue, of a nose, and of ears and eyes consummated
after the fashion of these organs in man; they are devoid
of an osseous, muscular, and myelonal (spinal chord)
system, and therefore of the nose "in toto," as being the
anterior extremity of the myelon.
3079. Such are what have been called the Invertebrate
animals, which are consequently, in accordance with
their physiological signification, Splanchnic or Tegumentary
animals.
3080. The tongue exhibits for the first time in Fishes
a resemblance to the human structure, while their nose,
ears, and eyes have not yet attained the latter grade of
perfection. To the nose are wanting the posterior nasal
foramina, to the ears the external auditory meatus, to the
eyes the palpebræ and power of motion.
3081. In the Reptiles the nose opens for the first time
into the mouth, and serves for the thorough passage of
air. It is thus developed as in Man, while to the ears
the external auditory meatus and cochlea are wanting,
the eyes being barely endowed with lids and motion.
3082. In Birds, for the first time, the external auditory
meatus, as well as the cochlea, is exhibited in its perfection,
while the eyes are scarcely gifted with motion, and
have only the inferior lid perfect; the tongue and nose,
with the limbs also, have again become retrograde in
character.
3083. For the first time, in the Mammalia, the eyes
are moveable and covered with two perfect lids, without
the other organs of sense having suffered degradation
through this completion of the eyes.
3084. Thus in respect to the Senses there are only
5 animal divisions of equal value or worth. They
should properly be called classes; but, as the lowest
division, from comprising within itself the viscera or the
vegetative systems, is very rich in contents; were we to
call these divisions classes, many inequalities in rank, and
hence also in the number of the orders and families,
would originate—
| 1. Dermatozoa | Invertebrata. |
| 2. Glossozoa | Pisces. |
| 3. Rhinozoa | Reptilia. |
| 4. Otozoa | Aves. |
| 5. Ophthalmozoa | Mammalia. |
3085. Now, with the sense of feeling, or the integument,
the sexual system is associated or conjoined, and
that indeed as the first or lowest development of the
tegumentary system. Nevertheless, the sexual system
divides into two groups, into the sexual organs, which
are impressed with a true sensorial signification, and
into their product, or the sexual juices, and the ovum or
fœtus. With these two divisions the development of the
integument proceeds "pari passu."
In the ovum the tegument and its contents are not as
yet separated. Both consist of a transparent mucous or
gelatinous mass, as is exemplified by the vitellus and
albumen; such is the case also in the Infusoria, Polyps,
and Acalephæ or Sea-nettles.
In the sexual organs, however, both parts separate into
membranous capsules or cysts, and glandular contents,
as in the roe or ovary; milt or testes; kidneys, with
furthermore the oviduct, penis, and urinary cyst.
The latter are sentient membranes. Such is the case
in the Bivalve Mollusca, and Snails.
In the next place the tegument becomes, for the first
time, a self-substantial organ of sensation by appearing
as an envelope of the body; the vesical form is then
repeated, and by this means the annular character originates
as a series of cysts arranged one behind the other;
it is thus a veritable skin, from which, finally, the members
sprout forth, as in Worms and Insects.
The Dermatozoa will accordingly range in three stages.
1. Blasto- or Oozoa.
2. Sexual animals.
3. Cutaneous "
An annulated tegument, cutis or true skin, appears for the
first time in the Worms, with here and there lateral filaments
and tentacula. True feet and antennæ appear
in the Crustacea or Crabs. Lastly, feet and wings in
the Insecta or Flies.
3086. The external sexual parts, especially the male,
first make their appearance, and that indeed with a very
striking amount of development, in the Snails, and in like
manner the body of the Bivalve Mollusca or Mussels has
become almost a complete mass of ova or roe. In the
Cuttle-fishes the first traces or rudiments of urinary
organs appear. The animals which belong to this group
are accordingly the Conchozoa or Shell-animals.
3087. Animals, which are directly resolvable into
sexual fluids, or that represent parts of the ovum, are
the gelatinous Infusoria, Polyps, and Acalephæ. Unto
this category belong the Protozoa or Mucus-animals.
3088. The complete subdivision of animals, according
to the organs of sense, would consequently stand thus:
| I. | Dermatozoa | Invertebrata. |
| 1. Oozoa | Protozoa. |
| 2. Glandular animals | Conchozoa. |
| 3. Cutaneous " | Ancyliozoa. |
| II. | Glossozoa | Pisces. |
| III. | Rhinozoa | Reptilia. |
| IV. | Otozoa | Aves. |
| V. | Ophthalmozoa | Thricozoa. |
3089. Unto these organs of sense the anatomical, or
internal parts, are subordinated, and range parallel to
them in a striking manner. The following is their order
of succession in accordance with that of their origin:
1. Intestinal system.
2. Vascular "
3. Respiratory "
4. Osseous "
5. Muscular "
6. Nervous "
3090. That the vegetative systems are correctly
arranged after this manner, is proved chiefly by their
order of development in the animal series.
3091. The animals, occupying the lowest grade, are
nothing but an intestine, which is in many instances
scarcely separated or distinct from the tegument; they
are devoid of vessels and branchiæ, and are barely provided
with self-substantial ovisacs. Their body consists
of one or two concentric cysts of an homogeneous and
transparent substance—Intestinal animals, Protozoa.
3092. When the intestine is freed from the mass of
the body, both then obtain the form and substance of tegumental
cysts, whereof the external being only an intestinal
tunic, thus represents the peritoneum. They are
now, however, united by means of a vascular system,
which is again surrounded by a cyst, and thus by a
pleura. Their body consists of three concentric cysts;
intestine, peritoneum, and pleura. It contains too a liver
and self-substantial sexual parts—Vascular animals,
Conchozoa.
If these cysts be repeated in the direction of the axis,
the tegument becomes an annulated skin. An annulose
animal is a multiplied cystic animal. Associated with
this character, the respiratory organs are gradually evolved
to form vascular plexuses, branchiæ, feet, tracheæ, and
wings, while the sexual parts are for the most part
separated—Ancyliozoa, Respiratory animals.
3093. The osseous system first appears in the Fishes,
along with imperfect, mostly tendonless, and only white
muscles, as also a myelon that is only developed into a
stunted encephalon, in which the brain-organs of the
Thricozoon are in great part wanting.
3094. Typical, or true muscles, provided with tendons,
and of a red colour, are first exhibited in the Reptilia.
3095. A complete nervous system, pretty similar to
that of the Thricozoa or Pilose animals, having a cerebrum
and cerebellum, with similarly distributed and
delicate nerves, is first displayed in Birds.
3096. According to the anatomical systems there are,
therefore, six divisions of animals:
| A. | Splanchnozoa. |
| 1. Intestinal animals | Protozoa. |
| 2. Vascular " | Conchozoa. |
| 3. Respiratory " | Ancyliozoa. |
| B. | Sarcozoa. |
| 4. Osseous animals | Pisces. |
| 5. Muscular " | Reptilia. |
| 6. Nervose " | Aves. |
3097. The Pilose animals first originate through
completion and combination of all the organs of sense.
They are Æsthetic or Sensorial animals.
3098. The arrangement of animals, according to the
organs of sense, coincides consequently with that derived
from the anatomical systems, and each animal division is
therefore determined by two principal organs, by a vegetative
and an animal. Every animal is at once a vegetable
and animal body, the inferior kinds being partly so,
but the highest or the Pilose animals entirely so, or in
every respect, i. e. in them are found all the anatomical
systems, and all the sexual and sensorial organs.
3099. The signification of animals is accordingly as
follows:
| I. Anatomical Systems. | II. Organs of Sense. |
| A. Vegetative Systems. | A. Tegumental Sense. |
| 1. Intestinal animals. | 1. Oozoa | Protozoa. |
| 2. Vascular " | 2. Glandular animals | Conchozoa. |
| 3. Respiratory " | 3. Cutaneous " | Ancyliozoa. |
| B. Animal Systems. | B. Cephalic Senses. |
| 4. Osseous animals. | 4. Glossozoa | Pisces. |
| 5. Muscular " | 5. Rhinozoa | Reptilia. |
| 6. Nervose " | 6. Otozoa | Aves. |
| 7. Sensorial " | 7. Ophthalmozoa | Thricozoa. |
The ovum divides into vitellus, and albumen, along
with the calcareous shell, and into germ or envelopes;
the intestine into pharynx or stomach, bowels and absorbent
vessels, thus:
| 1. Gastric animals | Vitelline animals | Infusoria. |
| 2. Intestinal " | Albuminous " | Polyps. |
| 3. Absorbent " | Involucral " | Acalephæ. |
The sexual parts divide into female, male, and urinary
organs; the vessels into veins, arteries, and hearts, thus:
| 1. Ovarial animals | Venous animals | Mussels. |
| 2. Orchitic " | Arterial " | Snails. |
| 3. Renal " | Cardiac " | Kracken. |
The annulated tegument divides into papillæ, feet, and
wings; the respiratory organs into tegumental network
or rete, into branchiæ and tracheæ, thus:
| 1. Papillary animals | Reticular animals | Worms. |
| 2. Pedal " | Branchial | Crabs. |
| 3. Alary " | Tracheal | Flies. |
When parallelized with the organs of plants, the following
remarkable relationships between them are rendered
apparent:
| 1. | Cells | Stomach | Vitellus | Infusoria. |
| 2. | Bark | Intestine | Albumen | Polyps. |
| 3. | Root | Absorbents | Envelopes | Acalephæ. |
| 4. | Ducts | Veins | Ovary | Mussels. |
| 5. | Liber | Arteries | Testes | Snails. |
| 6. | Stalk | Hearts | Kidneys | Kracken. |
| 7. | Tracheæ | Retia | Papillæ | Worms. |
| 8. | Wood | Branchiæ | Feet | Crabs. |
| 9. | Foliage | Lungs | Wings | Flies. |
| 10. | Seeds | Bones | Tongue | Fishes. |
| 11. | Pistil | Muscles | Nose | Reptiles. |
| 12. | Corolla | Nerves | Ears | Birds. |
| 13. | Fruit | Senses | Eyes | Thricozoa. |
A. DIVISION INTO PROVINCES.
3100. The animal body divides first of all into the
vegetative and animal. There will therefore be animals
in which the former, and others in which the latter,
systems predominate. The kingdom consequently separates
into a vegetative and into an animal province. The
vegetative parts are all tegumental developments, and
thus the creatures in whom they prevail are Splanchnic or
Visceral animals, but the animal parts are developments
of the flesh, and constitute the Sarcose animals.
First Province. Splanchnozoa.
3101. Unto the Splanchnic or Tegumental animals
are wanting bones, muscles with the nerves belonging to
them, and thus the neural axis or encephalon; they are
consequently devoid of bone, muscle, and brain, being in
a word asarcose or fleshless animals.
The tegument is, however, the general organ of sensation
or feeling; they are thus Sensitive animals.
3102. In them, developments only of the sense of
feeling can occur, such as sensitive papillæ, tentacula,
feet, and wings. All the remaining organs of sense can
only be exhibited as rudiments of a very feeble or stunted
character. They do not possess a true tongue, nose, ears,
and eyes, i. e. constructed after the type of these organs
in Man. The eyes only, from their being the sense of
the animal system proper, can assume a definite kind of
development.
3103. But, these organs of sense are the sensorial
organs of the head, or rather are the head itself; the
Tegumental animals are therefore devoid of a true head.
They only possess such an one, in so far as it is determined
by the tegument and nervous sense, by the mouth
and the eyes.
These animals are what have been called Invertebrata,
a name which is, however, defective, from its
indicating the absence of one part only or of a single
animal system, while the word flesh comprehends bones,
muscles, and nervous mass; they are asarcose animals.
But even this last appellation is not correct, because it
is negative. Their positive system, or that under which
they actually exist, is the tegument; so that the name
of Dermatozoa or Sensitive animals, is the only proper one.
As the tegument includes the viscera and forms therefore
the trunk alone, they may thus also be designated
Trunk-animals.
Second Province. Sarcozoa.
3104. As in the animal body, bones, muscles, and encephalic
system are suddenly associated with the tegumental
system; so also does a second series of animals,
having these systems, suddenly originate. Now, as the
first formation of the osseous system is the vertebra, so
all these animals have, as is generally understood, a vertebral
column, and are on that account indeed Vertebrata;
but, they are much more than this, and hence the
title is of too limited a character. Besides there are
among them animals, in which only the chorda dorsalis
is present, without the ring of the vertebral body being
formed around it.
Along with the animal systems the head is for the first
time developed with its organs of sense—Cephalic animals.
These animals have consequently, in addition to the
general sense of feeling, a true tongue, nose, ears, and
eyes—Cephalæsthetic animals.
B. ANIMAL CIRCLES.
3105. Animal Circles are representations of entire
anatomical systems as self-substantial bodies.
3106. Now, the vegetative body divides into three
principal systems, into the Intestinal, Vascular, and Respiratory
system, with their functions, or the digestive,
nutritive, and respiratory processes. There are therefore
Intestinal, Vascular, and Respiratory animals.
Circle I. Intestinal Animals.
3107. The intestinal system is the first form of body,
or that from which the other systems have not yet separated.
The body of these animals consists therefore of
the homogeneous primary mass—the animal protoplasma,
mucus or slime—Mucus-animals.
Now, the protoplasma is a hollow globule. The intestinal
system is therefore nothing else than the original cystic
form. Thus, there are Cystic animals like the Infusoria.
Again, cysts can increase in no other way, than by
dividing into their like, or engendering cysts within
themselves. The first kind of increase or multiplication
of species is thus fissiparous, or produced by division.
The newly-engendered cysts are to be compared with
the vitellus, and when they have attained perfection, to
the ovum. They are therefore Oozoa or Ovum-animals.
In these animals consequently there are no separated
sexual parts, namely, in addition to the vitelli, neither
testes nor renal organs, or at least only obscure indications
of them. In these Cystic animals the lowest feeling
only, that of the sex or general sensation, can occur.
The Oozoa or Cystic animals, when compared with the
plants, are the first flower that has been set free, or in
other words, a flower which no longer stands polarwise upon
a stem, because it is not developed in the differencing air,
but in the indifferent water. It may be said that nature,
having brought matters as far as the development of the
sexual parts, then quits or passes out of the vegetable world;
while these parts, or even the entire plant, requiring now
no longer the stem and root, become a root themselves,
and in behalf of this enter the water.
Animals, which have the form of flowers, are round
or radiiform. There are Radiata or Radiated animals.
3108. These Anthoidal or Flower-like animals are Infusoria,
Polyps, Acalephæ; being single or double concentric
cysts.
3109. We may regard the Flower-animals as the fundamental
mass of the sexual parts, which has attained
unto free motion. They are sex throughout, or nothing
but sex; it cannot therefore be said that they possess
sexual parts like the plants, but that they are sexual
parts. They are sexual parts that swim.
3110. Formerly, most of these Anthoidal animals were
actually taken for real plants, on account of their floral
and ramular form, and even their very substance; so
slightly withdrawn are they from the vegetable kingdom.
The whole difference between the two is effected by
the water. Could we transpose them into the air, they
would then become real plants.
3111. Now, as the vegetable flower is not a mere
sexual system, but is also stock or trunk; so also is the
animal flower at once an organ of digestion, respiration,
and nutrition. The lowest condition of these organs is,
however, only one of absorption, evaporation, and rigidification;
these processes will therefore be present also,
though only upon the lower stage—they are Intestinal
animals; for in a simple intestine the same processes
could occur, only within each other, whereas they are
mutually dissevered or set apart in intestine, lung, and
capillary system or parenchyma.
3112. The sexual parts are themselves viscera, or the
viscera themselves are sexual parts, just as the fungus is
at one and the same time root and seed-capsule. The
sexual parts themselves absorb, respire, and nourish.
The floral sac is not therefore a mere sexual sac, but
also an absorbent sac; upon a somewhat higher grade
it is even a digestive sac, the sac-wall itself being a respiratory
and nutritive paries or wall.
The sexual function has at once become an ingestive
function, tending unto nutrition, or the deglutition of the
food is itself an act of copulation.
The sexual capsule in these animal flowers can as well
be termed stomach as uterus, and its wall ovary as well
as branchia.
3113. As being of a sentient, mucous nature, they
are point-substance or nervous mass. The tentacula
are higher organized stamina, and thus occur as cilia,
surrounding the oral aperture or mouth, as in the Infusoria.
These palpi or feelers are, from being organs of
ingestion, both male penes as well as digits or tongues,
as in the Polyps. Their structure is still wholly tubular,
while their elongation appears to be for the most part
effected by injection with water—they are absorbents,
as in the Acalephæ. The Oozoon brings forth young in
the same cavity, in which it digests and by which it
respires, and impregnates itself with the same filaments,
whereby it seizes, swallows, and tastes its food.
In the bottom of the cavity of the Germ-animals, granules
develop, which are born or escape through the
floral opening—pharynx, and again become similar
beings. In others the granules also sprout forth attached
to the walls of the cavity, remain there some time
united with the parent animal, and thus completely represent
the mode of propagation in plants by means of
gemmules or buds. Among the Polyps and Acalephæ it
is well known that the ova issue from apertures near the
mouth; while in the Actiniæ this is stated to take place
from the stomach. The ovaria are, as is well known,
situated between the stomach and parietes of the body.
3114. The intestinal animals are an entire animal
organism, but only in the chaotic condition. They are
the fundamental tissue, the cellular system of the animal,
and the higher animals are only separated cells.
3115. The propagation is in every respect similar to
that of plants. Now, as the seeds are the whole plant
upon a small scale, so is the granule or ovum the entire
animal; it is liberated through the pharynx and continues
to grow merely by increasing in size. But if the
young animal protrude through the tegument, that is a
true gemmiparous propagation.
3116. Those Oozoa, which like plants can develop
buds, consist of several animals, and may be cut in pieces
like plants; when each piece becomes again an entire
animal.
3117. The Oozoa represent those products of nature
which are prior or antecedent to the animal world;
namely, first of all plants, and further still the inorganic
kingdom also, or the earth, since they have
originated in the water and can be as well developed
from the stones as the Lichens. There are therefore
Lithozoa or Stone-animals, and Phytozoa or Plant-animals,
among the Oozoa.
3118. Would we compare these animals with the
parts in plants, they then represent their cell-development,
or cells, bark and root. They are themselves either
vesicles, as the Infusoria, or barks, as the Corallines, or
a root-like fascicle of tubes, as the Acalephæ.
3119. In the animals, however, the cells have become
stomach, the bark intestine, the root absorbents. The
Oozoa represent therefore the aggregate of the intestinal
system, the primary mass of the animal body.
Their whole body is a digestive body, or parenchyma,
traversed in many species in all directions by tubes or
absorbents, as in the Acalephæ.
As yet no nervous filament, nor any muscular fibre,
&c. has been separated from their mass, as in like manner
a tegument has barely been freed or separated from the
intestine. These animals are nerveless and sine-tegumental,
precisely because they are wholly made up of nervous
mass and tegument.
Circle II. Vascular Animals.
3120. In the next place the digestive separates completely
from the tegumental function, and each constitutes
a function for itself, but which now being separated
from its fellow could no longer subsist. Between the
two therefore is formed the nutritive function in the
vascular system.
Now, the vascular system of the self-substantial intestine
is the liver. This organ will therefore for the first
time make its appearance in this animal circle—Hepatic
animals. Upon a higher stage, salivary glands are also
developed on the intestine, and will here likewise begin
for the first time to appear—Salivary animals, Snails.
3121. Through the separation of the viscera from the
remaining substance, this must necessarily remain behind
as a hollow cyst or tegument investing the former. The
true free tegumental formation is therefore by no means
accidental, but is necessarily bestowed with the viscera
in the course of animal development. This tegument is
peritoneum. There are animals which are invariably
bicystic, but consisting of concentric cysts—abdomens.
Around the ventral integument, however, the vascular
system also forms its tunic; this is the branchial
membrane or operculum—pleura, mantle. There
are tri-cystic animals—intestine, abdomen, and thorax
disposed concentrically around each other—Mussels.
Their body is therefore not articulated, but its parts are
still inserted within each other.
The Vascular animals are consequently multiplied
Protozoa; or Mucus-animals of the second power—Ovum2.
The sense of feeling ascends upon its second stage,
when its organ, having been freed from the mass of the
body, surrounds the viscera as a self-substantial tegument.
The feeling is then no longer one of a merely
general character, but is a definite perception of external
objects, a passive power of feeling.
3122. True muscles could not as yet originate in this
skin, and that for obvious reasons, although fibres are present;
for the former are to be brought under the signification
of arterial fibres.
3123. Cilia, when furnished with fibres by whose
agency they are rendered moveable and susceptible of
inversion, are called tentacula, which here occur under
every variety of form.
3124. Would we compare the Vascular animals with
the plants, they must then represent their ducts, liber,
and stalk. The heart is the stalk, the arteries liber, the
veins tubes or ducts. These animals have assumed also
upon the whole the cauline or cylindrical form—Cauline
animals.
3125. From their having obtained in addition to the
intestine only the vascular system, they are still governed
by the water, and live therefore for the greatest part in
this element. They have the first mode of respiration
and thus an aqueo-respiratory process—possess branchiæ
but not tracheal tubes.
3126. The sexual parts, which in the Germ-animals
were still coalesced for the most part with the body, here
become self-substantial through the separation of the
teguments, make their appearance as a repetition of the
digestive system under the condition of a free or separate
system, and are evolved into true ovaria and even
male parts—Sexual, Glandular animals.
3127. The first step towards the evolution of male
parts is, however, only half achieved. Only one testis
originates, while the other remains behind as ovarium—Androgynous
or Bisexual animals.
3128. These animals, which are characterized by the
vascular system, and by the first self-substantial or external
sexual parts, which indicate organs of sense, are the
Conchozoa, such as the Mussels, Snails, and Kracken.
Circle III. Respiratory Animals.
3129. When once the intestinal and vascular system,
through perfection of their individual parts, such as the
liver and branchia, and through separation of the sexual
parts, are completed; then the individualization of the
tegument steps into view, and it becomes an independent
respiratory system.
3130. Through the increased process of oxydation the
tegument hardens and is converted into horn. All
induration, however, only takes place in opposition to
soft places. The tegument therefore separates into hard
and soft rings—Annulate animals.
3131. The annulate tegument is a tracheal tube
wholly converted into a body. To distinguish it from the
general tegument it may be called the cutis or skin—Cutaneous
animals. The annular tegument may be regarded
as a series of cysts placed one behind the other. The
Annulate animals are therefore multiplied Malacozoa;
Mucus-animals of the third power—Ovum3.
The respiratory organs, being in their lowest condition,
will not as yet be freed from the tegument; the vessels
simply form a network or projecting filaments and lamellæ—reticular
branchiæ; as in the Worms. The tentacular
organs, from being yet soft and thus scarcely moveable,
are still very imperfect. Upon the lowest stage the
tegument or skin simply feels; in the next place papillæ,
and finally filaments, originate, especially about the
mouth—Cutaneous, Papillary and Filamentary animals.
3132. If the tegument, as being the original branchial
membrane, is converted into horn; then the branchiæ
cannot continue as retia, but must elongate above the
tegument into filaments, ramules or lamellæ.
With this, these elongated branchiæ separate into two
organs, one part of them becoming indurated in like
manner with the general tegument, and supporting the
other as gill. Horny branchial filaments, which contain
vessels, nerves, and fibres, are called feet—Pedal animals.
3133. The limbs or members of these animals are
simply hollow tegument, hollow hair, and are therefore
thoroughly different from the bones or the animal system.
The tegument thus hardens around the soft parts and
the viscera. A horny coat of mail originates, and thus
we have horny or mailed animals, in opposition to the
Malaco-or Conchozoa.
3134. Beneath the horn, however, there must still be
soft skin, and this becomes fibrous by the strong oxydation
which it undergoes. Fibrous fascicles are attached
to the coat of mail and to the hollow limbs, and are
consequently within the tubes.
3135. These fibrous bundles are not flesh, but a fibre-drawn
tegument, so that there are also no true muscles.
They must on that account too be numberless.
3136. The articulations or joints are external not
internal; they thus consist of tegumental tubes, not of
bones, abutting against each other, and are not surrounded
by flesh. Hence, like all the preceding groups
they also are devoid of flesh—asarcose animals like all the
preceding ones.
3137. In the branchiæ, however, it is only the cauline
portions which become horny, while their ramules continue
to perform the respiratory function. The branchiæ
therefore are appended to the extremity or roots of the
feet; or rather these latter form the branchial arches.
Annulate animals, having true or hornified and likewise
annulated feet, are called Crustacea or Crabs.
Thus, in these animals the sense of feeling obtains special
and moveable organs; they are Tactile animals.
Tactile organs are prolongations of tegument moved
by muscular fibres, which in some degree adapt themselves
by pressure unto the forms of objects, or have the
power of seizing and retaining them—such as feet,
antennæ, maxillæ, palpi.
3138. But if the tegument be entirely converted into
horn, and the respiratory vessels have in this way disappeared
within it, then the internal respiratory organs are
formed by inversion of that part of the tegument which
is between the rings, and through these openings the
air penetrates to the internal parts—stigmata, tracheæ.
3139. The tracheæ can first of all originate when the
respiratory process has attained its highest development,
or, in other words, when the creature breathes air.
3140. Lastly, in the air-breathing Ancyliozoa even the
external branchial lamellæ harden and are converted into
wings—Pterozoa, Insects or Flies.
The wings of Insects do not correspond to the wings
of birds; they are not feet, but pedal appendages or
branchiæ, and thus are no new or unknown organ.
Circle IV. Sarcose Animals.
3141. The second animal province may be regarded
as the fourth stage in the self-substantial development of
the anatomical systems, although, properly speaking, it
ranks as regards its value or worth upon a level with the
three former circles, and resolves itself directly into three
stages, which accord with its three systems. But since
these stages are at the same time also classes; they should,
for uniformity's sake, retain the latter name.
3142. The animal kingdom accordingly divides into
four great districts or circles.
| Circle I. | Intestinal Animals, | Oozoa—Protozoa. |
| II. | Vascular " | Sexual animals—Conchozoa. |
| III. | Respiratory " | Cutaneous " —Ancyliozoa. |
| IV. | Sarcose " | ; Cephalic " —Vertebrata. |
C. ANIMAL CLASSES.
3143. The animal Classes may be designated as the
self-substantial representation of a stage in the development
of an anatomical system or of the inferior organ of
sense, and in the Sarcose animals of these systems themselves
or of the cephalic senses.
3144. There are then as many classes as there are
stages of development or systems. Thus the intestinal
system separates into pharynx or stomach, intestine and
absorbents.
3145. The vascular system into veins, arteries, and
hearts.
3146. The respiratory system into branchial membrane
or skin, into branchiæ, and into lungs or tracheæ,
i. e. air-tubes. Taken in a strict sense, such divisions do
not constitute classes, as has been already remarked.
3147. It is only the animal systems, which do not
divide into several functions, but remaining upon a level
with each other, are simply repeated in the higher
organs of sense.
First Province. Splanchnozoa, Dermatozoa.
FIRST CIRCLE. INTESTINAL ANIMALS, OOZOA.
3148. The intestinal animals are nothing but depressed
cysts. They rank therefore upon the lowest stage of
development, and consist of mucus or granular nervous
mass—Protozoa, Gelatinose animals.
3149. It is an established fact that, with animals as well
as plants, the first function consists in imbibition or
absorption, and the body must consequently be an
absorbent cyst or a pharynx, which nevertheless takes up
the food in the same way that the tegumental lymphatic
vessels absorb. We can therefore style these animals
pharyngeal or gastric, although the name is not perfectly
correct—Infusoria. Then the intestine associates
with the pharynx or stomach, so that they are Intestinal
animals—Polyps or Corals.
Lastly the intestine sends out absorbents, and then
the animal consists of pharynx or stomach, intestine and
absorbents; such may be termed an Absorbent animal—Acalephæ.
The Intestinal animals therefore divide according to
the developmental stages into three Classes.
Class 1. Gastric, Vitelline Animals.
3150. The lowest animals commence with the water,
which has scarcely become mucus; they are nothing but
drops, vesicles, which swim about independently—Protozoa
or Primitive animals.
3151. The Protozoa correspond to the vitellus or the
male semen, which is nothing but vitellus in a state of
solution. They are the animal semen of the planet, the
animal dissolved. Animal generation cannot take a
deeper commencement or origin. The stone, which is
decomposed into carbon mixed with water, can become
nothing less than a point. They are the animal germinal
powder. The fungus is a tissue of vesicles, which
dissolves directly into seeds—fungus-powder or dust.
Thus, they are ovaria or testes which have dissolved into
seed, fluid testes—Vitelline, Seminal animals.
3152. The vitellus or semen is point-or nerve-mass
dissolved. The Vitelline animals are sentient or nerve-points,
which have combined all other processes in this
identical mass. The divided point-mass ranks, however,
in the signification of vesicular or cellular tissue. These
animals are nervose cells.
3153. Nerve-cells must originate in every water,
because every water is in a state of tension with the
earth and the air; thus, dissolves the former and absorbs
the latter. The water itself is a digesting and respiring
mucus.
3154. The nerve-cells have an internal cavity, from
the surface becoming oxydized and consequently converting
itself into a denser layer, or into tegument. This,
however, can only take place at the expense of the
internal mass, as being that alone which abuts against
the external parietes and becomes rigid.
3155. As the animal life is not simply a single act of
rigidification, but a repetition of the same, alternating
with solution, so must the Protozoon again replace the
mucus-granules which have been disposed of from its
interior; it must eat.
3156. It is a matter of indifference for the philosophy,
whether the reception of food is effected by one or several
mouths. There are Acalephæ and even Intestinal worms
(Entozoa), which absorb by several mouths, almost like
plants.
3157. But in the animal the mouth or mouths is or
are definite organic openings, not merely interspaces or
pores as in the plants; for they rank in the signification
of the blossoms, or of composite parts.
3158. It can be therefore said that every animal hath
a mouth or mouths, and consequently a stomach or
stomachs.
3159. Their motions consist in abbreviations or straitenings
of the cyst. Indications can scarcely be present
of the secretions of higher organs, such as the intestine,
vessel, branchiæ, liver, and such like parts.
3160. On the contrary, developments of the tegument
and nerves may occur, the former as cilia which serve also
as organs of motion and as branchiæ, the latter as ocular
points or rudimental eyes; for both are none other than
nervose tegument. The mouth of these animals is still
passive, and subordinated to the water. It is surrounded
only by cilia, which by the vortices produced by their
movements in the water, impel or draw the latter into
the mouth and with it the food. Such animals are called
Infusoria.
3161. As the Infusoria are the seed or vitellus itself,
so also are they the very ovum, and require no special
sexual parts for propagation. They absorb, and are so
nourished, and if there is a sufficient quantity of the mass
to admit again of its division into several points, they thus
divide. A magnified Infusorial animalcule has become,
as it were, an ovarium or a testis, which then produces
seed by dissolving itself into the latter. They are a
constant conflict of the organ and its product, of the
Solid and Fluid, a vitelline and orchitic process.
Class 2. Intestinal, Albuminous Animals.
3162. With the separation of the cyst into an internal
and external layer, or into intestine and tegument, the
animal must necessarily ascend a stage higher, since it
now contains two systems different from each other, and
is consequently a double Infusorium.
3163. The form of the Intestinal animals passes gradually
from that of the globe into the tube—Tubular
animals.
3164. They are tubular nerves surrounded by a
tegument.
The cilia also are perfected and lengthen into filaments,
which no longer perform simply vortical movements,
but now actually seize and convey the food
self-substantially into the mouth. Such animals are called
Polyps. Their multiplication takes place no longer
by division or fissure, but by ova and shoots, or ramification.
The oviducts or egg-tubes lie between the intestine
and tegument, and open upon the margin of the mouth
between the tentacula. In many instances the ovisacs,
or cysts, hang also freely from other parts of the body,
as in the Sertulariæ.
The sprouts or offshoots detach themselves from the
parent and become independent animals; but they frequently
continue to stand as ramules upon the maternal
animal, though they nourish themselves independently
of the latter.
3165. If the process of oxydation be augmented, then,
the tube's external wall indurates, becomes coriaceous,
and, lastly, ceratoid or horn-like in texture. The nerve-tubes
or the animal proper can now swim no longer,
since one kind only of motion is left it, namely, to protrude
itself out of, and then retract itself within, the
tube. It consequently falls to the ground, and while the
external mucus hardens, it clings to the former; such
are the Sessile or fixed Polyps.
3166. Sessile Polyps having coriaceous or horny tubes
are called Plant-animals, Zoophyta, Phytozoa.
3167. The adherent, dried and dead external tegument
of the Polyp is called stem. Such a ramified stem
completely resembles a plant.
3168. These woody or herbaceous stems are not rooted
in the earth, but have the power of adhering firmly to
every substance, to stone, glass, shells, and such like
bodies. They do not therefore draw in nourishment
through any root.
3169. The ramification is often wholly plant-like in
character, resembling that of a shrub with separate ramules,
which even assume too the form of leaves, and the
animal tubes that of flowers.
But frequently the ramules grow together also by their
extremities, giving rise to a trellis-work, the production of
which in plants is impossible. The soft animalcules,
which come in contact, cleave unto each other, and grow
together like wounded parts in the Sarcose animals.
3170. Upon the surface of the ramules or the leaves
are apertures, out of which the mucous substance protrudes
the radiated mouth. But these mouths are frequently,
especially in the Cystic corallines, of two different
modes of formation. The one kind are cysts without
filaments, and contain ova, which are developed and fall
out. The others have filaments, which move and do
not produce ova. The former look like seed-capsules,
the latter like flowers with stamina, while the entire
trunk resembles a monœcious plant.
3171. With increased oxydation calcareous earth is
deposited in the external tegument or rind, and the stem
is converted into stone—Lithozoa, Lithophyta, Corals.
This calcareous earth contains the most general acid, or
the carbonic, and thus oxygen combined with the inorganic
carbon, while the bones contain phosphoric acid,
oxydized gelatine.
3172. As the calcareous earth is, properly speaking,
only a granular deposit in the tegument, like it is in the
cartilages of the higher animals; so is it not to be regarded
as a free worm-tube, but as the body itself. Meanwhile
it forms a tube open above, from which the mouth of the
animal projects.
3173. As the animal ramifies, so also do the stony
tubes increase, and there originates a phytoidal or plant-like
stem, but one consisting of a stony mass.
3174. Thus the Coral is the earth-animal, and indicates
the globular or osseous mass under its first grade of formation
in the animal kingdom.
But there are also Polyps, whose stem only originates
through saccular inversion of the upper portion of the
animal's body; yet this is only distinct in the soft stems.
In most of the species, where a separate intestine is to be
found, it is probably only the upper part of the body which
is so inverted. Meanwhile there are some, whose intestine
forms a circle by returning upon itself, and opens into an
anus.
3175. If then the Infusoria are the vitellus or seed of
the animal kingdom; in like manner the Corals are its
ova. The carbonate of lime is the shell surrounding the
albumen, while the animal or intestine is the vitellus—Albumen-animals.
3176. Nature forms these living ova, when she takes
vitellus and albumen out of the sea-slime, invests them
with a shell derived from the earth, and hatches them,
after they have been vivified by sun and air.
Class 3. Absorbent, Involucral Animals.
3177. Did the former animals remain in the condition
of ova, from want of a perfect vascular system;
they are next developed, so soon as vessels appear and
form a vascular plexus, into fœtal involucra or envelopes.
These animals are vitellus with the vascular membrane.
3178. The Absorbent animals are no longer simple
vesicles, but large cysts or capsules like the developmental
envelopes of the fœtus, along with a choroid plexus—Involucral,
Fœtal-animals.
This choroid plexus does not, however, consist of arteries
and veins; but is only a ramification of the intestine, so
that the vessels are of a lacteal character—Absorbent
animals.
3179. In these animals there is no longer any egg-shell,
but everything has been taken up into the galvanic
circle; the shell has itself become organic and life-imbued.
Their substance is still mucous or albuminous;
they are still vitellus, though converted into a vascular
tissue.
3180. They therefore cling firmly nowhere; but swim
about freely, like brain-masses converted into radiated
cysts.
3181. Free Mucus-animals, traversed by vascular
plexuses, are Acalephæ.
3182. There are Acalephæ which are simply air-sacs,
like the air-sacs of ova, to which hang ramified vessels as
absorbent tubes—Cystic, Tubular Acalephæ.
Others represent hemispheres with numerous absorbent
tubes, which concur in the middle of the animal to form a
kind of stomach, from which again other tubes pass towards
the border, in order to elongate into tentacula. Thus
the absorbent vessels have become motor and sensitive
organs.
Besides this, many have around the mouth four large
lobes, which must be viewed as the antetypes of the sentient
lobes of the Bivalve Mollusca.
Lastly, others have a true mouth, which leads to a
similar gastric cavity, out of which the same vessels
emerge and ramify. Both kinds are called Hutquallen
or Acalephæ.
There are yet others having the same structure, but
oviform, with respiratory lamellæ upon the absorbent
tubes, Rippenquallen or Beroes.
3183. An Acalephan is a brood-egg, which swims
freely about without a shell.
3184. The vessels are quaternary in their distribution,
and form a cross, like the involucral vessels of the chick.
3185. In these animals the ova first begin to be detached
and agglomerated together in definite situations
so as to constitute ovaria. The number here is also four.
They lie usually within four cavities surrounding the
stomach, and into these wide apertures situated near to
the mouth lead; they are at the same time regarded as
respiratory cavities.
In other species vesicles, wherein seminal animalcules
are developed, occupy the same situation. Here therefore
is recognised for the first time a separated sex.
In the Röhrenquallen or Physaliæ the ovisacs mostly
depend externally in the form of bells.
Besides this we find in the Acalephæ all sorts of laminæ,
which are probably organs of respiration.
It is not known what is the meaning of the air-sacs
and of what the air consists.
Most of the species emit light like globes of fire, just
as many Infusoria do also. It is probably a phosphorescence
given out by the mucus when passing over into
a state of decomposition.
Very many have also the stinging property of the
nettle; but, whether the cause of this is chemical or mechanical,
is not yet exactly known.
SECOND CIRCLE. VASCULAR, SEXUAL ANIMALS.
3186. So far, or in the ascending scale up to the Acalephæ,
the animal is only viscus with an absorbent
canal, which is at the same time a canal of evacuation,
without a distinct intestine being set apart for that purpose;
such is the general rule.
3187. After the Acalephæ the formation undergoes a
change; the distinction between the exterior and interior
is prominently displayed, and the internal wall becomes
freed as a separate and perfect intestine along with a
mouth and anus; the external wall appears as a free
tegument. But, seeing that two concentric and separated
cysts could not subsist without combination by means of
the nutritive system or the vessels; a perfect vascular
system is formed, divided into veins, arteries, and hearts.
The tegument, wherein the vessels become self-substantial,
is the branchial membrane. There is therefore placed
around the intestinal body a vascular body also, or
branchial membrane, which is consequently pleura or
mantle (pallium), as in the Mussels. The body of the
intestine consists of intestine and peritoneum; that of
the vessels of branchiæ and pleura or mantle.
These animals are hence bisystemic, being both Intestinal
and Vascular animals; but, since the vascular
system is a new addition, it is thus characteristic of the
circle, and its members must be therefore called Vascular
animals. With the vascular system, however, all its further
developments have been bestowed; thus especially
the complication of the vessels with the intestinal ramifications
or, in other words, the liver—Hepatic animals.
The salivary glands also are a similar complication, and
in this series therefore they make their appearance.
Lastly, the renal glands are such a kind of vascular
organ, or branchiæ of the sexual parts; they also begin
to be astir in this series.
With the separation of the systems into separate
teguments or membranes, the sexual parts also separate.
The ovarium becomes an independent organ furnished
with its own excretory ducts; the male parts are individualized
to form veritable testes furnished with excretory
ducts, or even with a penis. Still this is all
effected only by degrees, or as yet within the confines
of this circle.
These animals divide according to the viscera into
Venous, Arterial, and Cardiac animals; according to the
sexual parts, into Ovarial, Orchitic, and Renal animals.
Class 4. Venous, Ovarial Animals.
3188. With the protrusion into view of the vascular
system, the veins are chiefly developed along with their
principal organ, the liver, as being the bond of union
between the circulation and the intestine.
3189. The animals, which first bring to bear in addition
to the intestine a liver, are the Bivalve Mollusca.
3190. With the veins arteries also originate, but with
a preponderance of venosity or the venous character. The
blood is lymphatic, colourless.
3191. The cardinal venous organ or the liver evokes
also into existence a corresponding organ of respiration,
namely, free branchiæ with the tegumental form—branchial
laminæ or leaves.
3192. In the middle between the branchial laminæ
and the liver, the heart is evolved; this organ comprising
always a ventricle with an auricle, but consisting of a
venous, membranous tissue almost devoid of fibres.
3193. The first heart is in other respects arteriose; it
receives the blood from the branchiæ and transmits it to
the liver as well as to the remaining parts of the body,
from which it proceeds directly to the branchiæ without
entering any venous or right heart.
3194. In these animals, as is well known, four branchial
leaves lie externally on the belly, which includes
the intestine with a large liver, and hangs together with
the branchiæ as a separate purse within the mantle.
3195. In the Mussel a structure originates for the
first time, which can be compared with a thoracic or
pectoral cavity. What covers the branchiæ, must stand
in the signification of the thorax or chest. The pallium
or mantle of the Mussels is pleura.
3196. Their shells are branchial opercula (as in the
Fishes). They are secretions from the mantle, and everywhere
accompany the branchiæ.
3197. The locket or hinge of the shell-valves corresponds
to the rachis or spina dorsi, as is especially distinct
in the Teredines or "ship-worms."
The shells of Bivalve Mollusca are a calcareo-thoracic
box, open in front, inverted behind, and moveable like ribs.
3198. The two sphincter or occlusor muscles signify
shoulder and haunch.
3199. These animals begin for the first time to exhibit
bilaterality or symmetry, because in them there is stirring
the idea of the osseous formation. From the branchiæ
being situated symmetrically upon both sides, the cardiac
chambers are also symmetrical.
3200. The pectoral tunic (mantle) usually elongates at
the anal extremity into two tubes, respiratory tubes,
through which the water is drawn in and thrust out or
expelled. Such is the case in the highest Worms or Holothuriæ,
only there the respiratory tube leads into the body
itself. A similar arrangement is found in the Echini.
Many aquatic larvæ from all kinds of Insects or out of
different classes breathe through anal tubes. All these
animals consequently repeat the Mussels, and this formation
admits of being followed out up to Man, where it is
left as allantois and primordial kidneys. The thorax of
the Mussel thus opens by the anus. But, since the thorax
is that which is here preponderating and is nearly the
whole animal, so does the anus open into the posterior
respiratory tube.
3201. The Mussel can be regarded as an animal consisting
of three cysts or sacs inclosed within each other,
namely, of intestine, around this the ventral pouch, and
around this again the thoracic sac or mantle. If, moreover,
the shells be regarded as a cyst, the animal then
consists of four sacs. The heart and branchiæ lie within
the thoracic cavity; intestine, liver, and ovarium within
the abdominal or ventral cavity. The Mussel is thus a
doubled Acalephan.
What is termed foot in the Bivalve Mollusca is nothing
else than the abdominal tegument dilated in front to
form a muscular ridge.
3202. The Mussels are embryos, in whom the liver
originates, and whose chorion acquires a placenta. Then
again, as the embryo is nearly all liver, and hangs surrounded
by a watery fluid within the widely-expanded
chorion and amnion, so does the abdominal pouch hang
within the pallial cavity, or in the wide, water-full chest.
3203. In the abdomen there is still only the ovarium,
and that indeed very large. There are properly two
ovaria, each of which, according to my observations (vid.
Göttinger Gelehrte Anzeigen, 1806. Stück 148), opens
laterally under the shoulder-muscle and gives exit to the
ova, which then enter within or between the branchial
fringes, in order to be there developed.
3204. Here the respiratory organs are still at the
same time a kind of uterus. The ova may be oxydized
therein, like the embryo is in the uterus. These branchiæ
are probably still to be compared with, or designated
as, sexual branchiæ.
3205. In the back of the shoulder these animals have
a highly vascular organ with two excretory passages,
which open near to the orifices of the oviducts. I formerly
regarded it as a kidney, but, according to more
recent observations, it should be a testis. In the anterior
part of the foot there is frequently situated a gland
which ejects a gelatinoid moisture, which hardens into
glutinous filaments—the byssus or beard. It is probably
a memento of the tentacula of the Acalephæ. There
is besides also an organ in the feet of many Bivalve
Mollusca, which occasionally squirts a watery juice out
to a great distance. I have occasionally found this organ
in our Teichmuschel. Its signification is probably the
same as that of the preceding organ.
3206. The organization of the Bivalve Mollusc can be
thus most distinctly described; it is an abdomen, wherein
is an intestine with mouth and anus, a liver and a double
ovarium; upon the sides of this abdomen are situated
the branchiæ in the form of four laminæ; around the
branchiæ and the abdomen is the pleura or mantle, which
is always open posteriorly.
3207. The mouth is placed directly at the anterior
extremity of the abdomen, is devoid of neck and head,
as also salivary glands; it is consequently not a true
mouth, but only a pharyngeal aperture. Thereupon,
however, are situated four sensitive lobes, which are in
structure exactly like the branchiæ—cephalic or pharyngeal
branchiæ. They are the further formations of the
four arms of the Acalephæ.
3208. The Mussel has a perfect splanchno-neural system
with ganglia and a pharyngeal ring, which probably
corresponds to the nerves that sweep around it.
3209. The Mussel has no other organ of sense than
that of the passive sense of feeling, the tegument. It
cannot once move its sensitive lobes voluntarily; it has
no lips.
3210. The abdomen only elongates in most of them
to form a moveable, variously constructed foot or keel,
which cannot, however, creep but only push. The progression
of the Bivalve Mollusca is backwards, as in the
Acalephæ. In the Snails the ventral surface first becomes
a creeping sole.
3211. The Mussels repeat the Infusoria; are Infusoria
with a bivalve calcareous testa or shell.
Class 5. Arterial, Orchitic Animals.
3212. In the preceding class of Bivalve Mollusca,
it is in truth the abdominal viscera only, such as the
intestine, liver and ovarium, which have been perfected;
and then the veins and arteries with a membranous
heart. The cephalic organs, eyes, maxillæ, salivary
glands, and even moveable lips with tentacula, are wanting,
as well as the muscular heart. Lastly, the arterial
sexual system or a self-substantial testis and the penis
were absent.
Now, Mollusca, which have eyes, maxillæ, a muscular
heart and a ventral sole or foot, with salivary glands and
a penis, are Gasteropoda or Snails.
3213. The Snails possess salivary glands, a trace of the
tongue and of maxillæ, moveable lips and tentacula, with
thus an approximation to the head, unto which the eyes
are rarely wanting—Salivary animals.
3214. With the development of the head or rather of
its inferior organs of sense, the antagonism makes its
appearance also in the ovarium. A half of the ovarium
is converted into the testis.
The Snail is therefore a Mollusc, which is female upon
one side, male upon the other.
3215. The androgynous or bisexual animal is, as a
general rule, asymmetrical.
3216. The mantle also or the branchial cavity obeys
this want of symmetry. The branchiæ of one side dwindle
down; those of the other turn with the mantle towards
the head, and the respiratory aperture occurs upon the
back.
3217. With the one-sided evolution of the mantle,
one shell also is only developed, while the other is
stunted or placed under arrest. The Snail's shell is one
of the Bivalve Mollusc's shells, its operculum is the
other. This last is stony, horny, and finally is entirely
wanting.
It is remarkable that the right shell has been pretty
generally perfected, while the left dwindles down into the
operculum; all the Snail's openings are therefore upon
the right side, such as the anus, with the orifices for
the escape of the ova and semen.
Male animals are situated on the right side, female
on the left; or where the right side was predominating,
there originated the male sex, where the left, the female.
3218. As the orifice of the mantle and the shell is
properly the aperture of the branchial foramina; so can
it be said, that the Snail were a Mussel, which does not
simply extend the foot, but also the mouth or head to
the branchial opening; it is a Mussel reversed.
3219. Every thing else in the Snails is arranged in
accordance with these fundamental organs and forms.
The cephalic portion of the intestinal canal is indicated
also by muscular fibres. The pharynx and the mouth
can contract and expand, seize and bite off; the former
frequently admits of being protruded and retracted as a
fleshy proboscis with perforating maxillæ.
3220. Since the muscular fibres are only tegumental
fibres, and therefore lie within cavities; they thus act
like as in the feet of Insects.
The tentacula of many Snails are moved like the feet
of Insects; but as they are not horny, but soft, they are
turned either inside out or "vice versâ." The oviduct
and seminal duct or penis follow the same mode of formation.
They are likewise everted and inverted.
3221. These members of the Snail are true Insect-limbs
that have remained soft, and are thereupon susceptible
of inversion and eversion. Were an Insect's foot
soft, every one will admit that it would then yield so as
to become inverted, if the fibres pulled upon it. The
limbs of Insects have thus only become stiff, and are
thereby Snail's horns that resist inversion. All these members
are teguments, and give the lie only unto limbs;
for it belongs to the essence of a limb, that it be dense.
3222. That which suffers eversion or its converse is
no limb, but only a sheath, a prepuce. Nearly the whole
Snail is but a prepuce, a "membrum virile" or "verge."
3223. There is no class of animals, in which the testes
and penis are found so disproportionately developed as
in the Snails—Orchitic, Penes-animals.
3224. The vascular and nervous systems are related
pretty nearly as in the Mussels. But the heart is fleshy
and has, by reason of its unilateral or single branchia,
only one auricle also.
3225. The Snails repeat the Corals in the cylindrical
form of the body, the tubular-shaped shell, and the return
of the intestine upon itself towards the mouth, as in
many Corallines.
3226. In them also the organ, which virtually corresponds
to the kidney, appears to be astir, namely, what has
been called the calcareous gland, situated within the
branchial cavity, and which opens not far from the anus.
Class 6. Cardiac, Nephritic Animals.
3227. Hitherto there has been found only a single
"cardia," namely, the left or arterial heart, which, receiving
the oxygenated blood from the branchiæ, propels
it to all parts of the body—for the purposes of nutrition.
But now also the right heart suddenly makes its
appearance, which drives the blood into the branchiæ—for
oxygenation. This must doubtless be regarded as a
higher development, especially since, as I have shown, in
the higher animals as well as in the embryo, the right
heart is first perfected subsequently to the left.
The Conchozoa with a double heart are the Sepiæ or
Cuttle-fishes. These are in consequence correctly called
Cardiac animals, and must be regarded as the fundamental
form of this stage.
With this completion of the heart other structural
changes, which probably stand in intimate connexion
with it, appear. In the Snails indeed a kind of renal
organ, or what has been called the calcareous gland
in the branchial cavity, has been already shown to
exist. Whether the shoulder-gland in the other Mollusca
belongs also to the members of the present class,
may be left undetermined. This kidney, like other
vegetative organs, pours out its secretion quite involuntarily.
In the Cuttle-fishes it is, however, combined with
an organ, by means of which it can discharge its fluid,
or the ink, voluntarily as in the higher animals. The
kidney or ink-gland with the ink-sac is therefore characteristic
likewise of the Sepiæ, and admits of our calling
these creatures Nephritic or Renal animals.
At the same time the whole form of the body is altered;
it becomes cylindrical, and is provided upon the ventral
aspect with neither a muscular keel nor sole, whereby it
might glide or shove itself along; thus herein also a
resemblance is manifested to the higher animals.
This motionless body is on the contrary endowed
with independent organs of locomotion, namely, with
fins or arms, which are wanting in the Snails and
Bivalve Mollusca. The labour of motion is consequently
removed by the organs which minister unto it from the
body, and thus we trace another similitude to the higher
animals, which transport their body from place to place
by wings or feet. These animals can be therefore called
Cylindrical Snails, out of contrast to the Sole and Carinate
Snails or the Bivalve Mollusca.
Closely allied now to these cylindrimorphous Conchozoa
are all those Mollusca in whom the keel (carina) or sole
is wanting; i. e. such snail or mussel-like animals, which
either cannot transport themselves at all, or effect this
by means of fins or podoidal appendages, in short all
that have been called Snails, having no sole, and all
supposed to be Mussels, but without a keel.
Now, the Heteropoda or Pterotracheæ and the Pteropoda
or Clionæ have projecting processes which act like
fins.
The Brachiopoda or Terebratulæ, though unendowed
with the power of locomotion, have, as their first name
implies, arm-like organs.
The Cirripedia or Lepades, being likewise without the
power of continuous motion, possess podoidal or foot-like
appendages. In both these families, however, these
appendages are nevertheless true motor organs; for by
means of them they seize their food, an act which
hitherto has been seen to take place neither in the Snails
nor Bivalve Mollusca.
Lastly, the prehensile organs dwindle down into mere
filaments or small lobes, but the body always retains its
cylindrical form without keel and sole, as in the Ascidiæ
or Meerscheiden.
In the Salpæ only the cylindrical body is left, unto
which appendages are still, and not rarely either, attached,
serving them wherewith to lay hold of each other. In
accordance consequently with their external form all these
animals appertain to one class along with the Cuttle-fishes.
I call them Kracken.
The renal organs are still not to be found in all of
them; the structure alone of the heart agrees essentially,
despite of its simplicity, with that of the Cuttle-fishes.
In the Ascidiæ, simple as well as compound
species, it is indeed but a single, though muscular
pouch. Now, this pouch drives the blood alternately, at
one time into the branchiæ, at another backwards into
the body, and is thus in the first act a venous or right
heart, in the latter an arteriose or left; consequently,
according to function this single pouch is compounded or
made up of the two hearts.
The branchiæ deviate entirely from those of the
Mussels and Snails, and exhibit a very complicated structure;
in the mussel-like families they present the form
of a trellis-shaped sac, as in the Ascidiæ, or that of filiform
appendages to the feet, as in the Cirripedia, or they
are funiform as in the Brachiopoda. In the Pteropoda
they are very varied; in the snail-like Heteropoda they
are mostly pectiniform or tuft-shaped; in the Sepiæ or
Cephalopoda phylliform or fin-shaped.
The external character ccommon to the whole class is
the cylindrical body, unto which may be aptly conjoined
the possession of special organs of motion; whether
such organs be prehensile in function, media of support,
or veritable instruments of progression.
The snail-like Pteropoda, Heteropoda, and Cephalopoda,
have a head, but this is wanting in the mussel-like
Salpæ, Ascidiæ, Cirripedia, and Brachiopoda.
THIRD CIRCLE. RESPIRATORY, CUTANEOUS ANIMALS.
3228. Respiratory animals are Dermatozoa with a
predominating system of respiration.
Now, the respiratory system is the tegument, which here
then must attain the highest pitch of perfection.
This again takes place through the increased process of
oxydation, which produces induration of the parts.
The vessels, which surround the tegument, must
nourish this portion of it to a greater degree and render
it harder or more compact than any other, whereby
alternating expansions and contractions thereof originate,
in other words, the structure of the trachea.
3229. The whole body of the Respiratory animals
becomes a trachea, a series of rings. The Respiratory
are therefore the Annulate animals.
The rings are to be regarded as cysts, which are
mutually juxtaposited or repeated, not being inserted
within each other as in the preceding animals, but behind
each other, so that the present circle consists of multiplied
Cystic animals.
3230. The Annulate animals must represent the
developmental stages of the tegument, while the viscera
make a retrograde step—Tactile animals.
3231. The tegument has two functions: it is an
organ of respiration and of general feeling or sensation.
In both cases it passes through three stages of development.
It is either entirely a branchia, i. e. reticular in character;
or the branchiæ become individualized in certain
situations and partly horny, constituting branchiæ proper;
or finally, the tegument is converted into an air-breathing
organ, tracheæ or air-tubes.
3232. It either feels with its whole surface, or by
papillæ; or the branchiæ change into horny feet; or
lastly, into alary appendages or wings. The Ancyliozoa
consequently divide into Reticular or Papillary animals;
into Branchial or Pedal, and into Tracheal or Winged
animals.
3233. As the arterial character preponderates in these
animals, so do the venous organs, especially the liver
and renal structures, retrograde.
3234. The body is now an intestinal and tegumental
body with predominating respiratory and sentient organs;
here therefore sensitive papillæ, antennæ, feet and wings,
appear in abundance.
3235. With the retrogression of the viscera, the
glandular structure also and large proportional bulk of
the sexual parts disappears. They assume the form of
the intestine and tegument, i. e. become tubes.
The spawn like the milt consists, as a general rule,
of only two long membranous tubes, running near the
intestine.
3236. The external sexual parts belong to the sense
of feeling and follow the developments of the tegument;
they make their appearance as antennæ, and are usually
accompanied by foot-like accessory organs.
3237. The nervous system in like manner accompanies
the tegument. It consists of two gangliated
cords passing along the ventral surface.
3238. As the maxillæ are only repeated feet, so they
are rarely wanting, and are, like the latter, disposed after
the fashion of scissor-blades. The same holds good of
the antennæ; they are mostly in pairs, and consist of a
series of rings projecting from the head.
3239. The eyes are usually present, but only as tegumental
organs, or placed upon the points of the antennæ.
Class 7. Reticular or Papillary Animals.
3240. The repetition of the Intestinal or Vascular
animal, taking place under the dominion of the respiratory
system, must be accompanied by the vascular
system. The tegument is a vascular, a branchial membrane,
a branchial network or skin.
3241. A body with predominating tegumental and
intestinal system, is cylindrical. It can only become
gross or lump-shaped, if the glandular viscera, such as
liver, ovarium and testes, get the upper hand or prevail.
3242. A tegument which appears as a branchial
membrane, is annulate or ringed.
3243. An animal, having an annulate respiratory
membrane without annulated feet, is a Worm.
Upon the lowest stage the vascular system is still
similar to that of the Vascular animals; its blood is a
colourless fluid—White Worms, Entozoa.
In order to be a Worm, it is sufficient to have an
annulate tegument. Even if the intestine be wanting
and the tegument supplies its place; the character is
nevertheless complete. Here it is the skin, which digests,
while in the Intestinal animals it is the intestine, which
digests and breathes. Since it is here the tegument,
which undertakes the offices of the whole body; the
vessels, liver, and salivary glands are wanting unto the
intestine.
The sexual parts are also frequently arrested and the
ova appear to originate in the tegument.
Many undergo division without injury to themselves;
yet the detached portions do not become again entire
animals.
In most of them, however, the sexual parts of both
sexes are present, being associated with, though separate
and distinct from, each other; those of the female mostly
opening upon the sides of the body.
As being Dermatozoa, they have for the most part
special organs of sensation around the mouth, under the
form of papillæ, filaments, spines and cups. The nervous
system, wherever it is to be met with, consists of a ring
surrounding the pharynx, and of a double ganglionic
cord lying along the ventral surface of the body.
3244. Upon a higher stage of development the arterial
system gains the preponderance, and the blood
becomes mostly of a red colour—Red-blooded Worms.
3245. With the arterial the fibrous system also makes
a more decisive appearance. The tegument is a fibrous
tunic—it is itself an artery. All Annulate animals with
a fibrous tunic, which can consequently contract, belong
to this class, whether too they have red blood or not,
like the Holothuriæ and Star-fishes.
3246. The Lumbricales or Earth-worms and the
Hirudines or Leeches respire obviously through the whole
tegument, though a special organ of respiration does
begin to be evolved, in the first family in the "saddle," in
the second in the lateral vesicles or cysts.
3247. In others the branchial vessels make their
appearance above the level of the tegument as filaments
or ramules, and are arranged in two rows, as in
the common Sea-worm and in the Nereides.
3248. Lastly, they accumulate about the neck or
head, as in the Amphitrites and Serpulariæ.
3249. There are also Worms which respire only
through the intestine, its vascular system being bathed
on all sides by water, as in the Aphroditæ. This water
becomes, as it would appear, simply imbibed by the
tegument in Thalassema, but through an opening at the
anal extremity of the body in the Holothuriæ.
3250. These Worms have also no liver, or at the very
best only traces of that organ, by reason of the preponderance
of the arteries.
3251. Papillæ or sensitive filaments gradually sprout
from the external branchial filaments, lie along the sides
of the body, and are the prelude of the feet; as in the
Nereides—Papillary, Filamentary animals.
3252. Others of these filaments become horny and
appear as setæ or bristles, placed somewhat similarly to
the above, as in the Earth-worm.
3253. Mouth and head are more perfectly developed
than in the Entozoa. The first can readily expand and
contract, frequently protrude the pharynx like a proboscis
or tube, and not unfrequently has manducatory
pincers, like as in Insects.
3254. On the head there are mostly annulate tentacula
with muscular fibres, and frequently simple eyes.
3255. In those species which draw the water itself
into the body and use it for the purposes of respiration,
the formation of the mouth has risen higher, and the
pharynx been provided with maxillæ, in number five
or ten—Echini, Holothuriæ. These maxillæ form in
themselves a peculiar skeleton around the pharynx, which
is ranged circularly instead of by pairs.
3256. The nervous system is directed according to
the relations of the tegument and intestine. It forms
two ganglionic cords running along the ventral side of
the body, and, where it makes with the nerves going to the
maxillæ a ring around the pharynx, corresponds to the
pharyngeal or pneumogastric nerves.
3257. In reference to the sense of feeling it may be
said, that the Worms were those among the Sensitive
animals, which feel by the whole tegument or body.
Their body itself is a tentaculum.
3258. The sexual parts are likewise intestini-and
dermiform, not glandular in figure like the ovarium and
testis of Mussels and Snails, but tubular, as in the
Entozoa. There are usually two oviducts and two
seminal tubes to be found.
3259. So far as we are acquainted with the Red-blooded
worms, they are androgynous, at least as regards
the Earth-worms and Leeches, and their sexual parts are
indeed tolerably symmetrical; yet they do not open
posteriorly, but far forwards on the ventral side of the
body, this being the case with even the male sex-organs.
Class 8. Branchial, Pedal Animals.
3260. The representation of the Cutaneous animals
is not yet the entire completion of the tegument. It
attains a higher grade by separating in accordance with
the visceral systems, and in the conversion of the branchiæ
into sensorial organs or feet.
The branchiæ endeavour to separate self-substantially,
and form a body of their own. The branchial body is
the thorax.
By this attempt, the abdomen originates "per se" as the
body of the intestinal and sexual system. The branchiæ
upon it are arrested or stunted. In like manner
the body of the nerves separates itself from the thorax
and becomes head.
But meanwhile this separation is only imperfect; the
head and thorax, if otherwise distinguished by their size
and the feet, are for the most part connate; abdomen
and tail are usually fused together, and only to be
recognized as distinct by the limbs or members.
With this greater division of the body each division
of it obtains its articulation into rings; but they will be
hardened by virtue of the separation of the branchial network,
or will become horny—Horny animals.
3261. With the conversion into horny texture of the
body, its branchial filaments or tentacula must undergo a
similar change, and divide likewise into rings like the
general or bodily tegument.
Annulate, horny branchial filaments, are feet—Pedal
animals.
A part of the branchiæ remains hanging to the feet,
mostly to their basal joints, and is supported by animal
motor members or limbs; the respiration becomes voluntary—Branchial
animals.
The Annulate animals, having branchiæ and feet, are
the Crustacea or Crabs.
The branchiæ and feet are, of course, most largely
developed upon the thorax.
Those upon the abdomen continue small, and those
upon the tail or sexual body dwindle in size and change
into other organs, such as fins, sacs, filaments, pincers.
3262. The number five of the branchiæ is also exhibited
in the feet. There are mostly five large pairs of
feet on the thorax, and as many small ones on the abdomen,
as in the Crabs.
On the tail they appear more dwindled in size, and
are frequently reduced to a smaller number.
Each part of the body has properly five rings; and
these parts being the thorax, abdomen and tail, there are
thus fifteen rings in all.
In the Crabs it is distinctly demonstrated that the
maxillæ are nothing else but feet; there are therefore
for the most part, after taking into account sundry arrests
and coalescences, five pairs of maxillæ also.
The Crabs have usually two pairs of antennæ, whose
signification is unknown; one pair is probably the elongation
of the ear.
All Branchial animals have eyes, frequently too supported
upon articulated pedicles.
They have a double nervous cord upon the ventral
surface of the body.
They have, like the Mussels, a heart upon the dorsal
aspect, and arteries with veins.
The intestine opens into the apex of the tail, and is
surrounded by a liver.
The sexual parts still open for the most part on the
thorax, and indeed by two orifices, as in the Worms.
There are no more androgynous or bisexual beings in
the present class.
In some few the branchiæ already enter the body and
become air-tubes, as in the Scolopendræ, Spiders and
Scorpions.
The Branchial animals repeat, as being the second
class of their circle, the Corals and Snails; their "quasi"
coat of mail is therefore harder, often richer in calcareous
ingredients, and, in addition to this, lies frequently
upon the thorax or branchia as a special testa, called the
scute or shield.
Class 9. Tracheal, Alary Animals.
Annulate animals, whose branchiæ have undergone a
partial conversion into tracheæ and into wings, are the
Insecta proper or Flies.
3263. A Worm with feet, tracheæ and wings, is an
Entomon or Insect.
3264. The first separation takes place in respect to
the three tegumental segments of the body, the abdomen,
thorax, and head. All three are, in the Insects,
more separated from each other than in the Branchial
animals, and united together usually by a narrow tube;
even in cases also, where they are connate with each
other, they are still easily recognized by respect being
had to size, form, or appendages.
Every Insect is divided into three segments. In the
abdomen are the organs belonging to the Worm, such
as the intestine, and a fatty body which appears to be
an analogue of the liver, a dorsal vessel, tubular sexual
parts and air-tubes (tracheæ), but nothing else.
The abdominal feet now disappear entirely, and even
the number of thoracic feet diminishes, owing, doubtless,
to the production of wings.
3265. The thorax alone is reserved for the limbs or
locomotive members. It never carries more than three
pairs of legs and two pairs of wings. Of the viscera, it
contains nothing but the pharynx, while in the Crabs,
important intestinal organs and even the liver, are situated
within its cavity. It is in Insects therefore nothing
else than a medium of support to the respiratory organs
which have become limbs. Hence the thorax never has
more than three rings, namely, one for each pair of
legs. The wings invariably stand upon its two posterior
rings.
3266. Since the limbs of the Insect are only the lateral
filaments of the Worms, which have become hard and
consequently hollow; they are as such not to be termed
true feet, but are only to be compared with branchial
arches or ribs; a step by which their greater number
also admits of being understood. They are not to be
compared with our feet, but unto the toes, which have
been separated from each other as far as the rings of the
body. The Crab has properly five thoracic and five
abdominal toes. All its thoracic feet taken together
are only equivalent to our hand.
The feet of the higher animals are only Insect-feet
connate or coalesced.
3267. In other respects they already typify or prefigure
true limbs, as well from their position as by the
division of their joints. A perfect beetle's foot divides
exactly like the limb of Man, into femur, patella, tibia,
tarsus, and phalanges. These parts of the leg must not,
however, be so absurdly divided and named, as has
hitherto been unfortunately the case in our systems,
where the femur has been called coxa, the patella trochanter,
the tibia femur, while the toes have been lumped
under the name of tarsus. (Ed. 1st, 1811. § 3087.)
The regular number of the toes or tarsal joints is five,
so that they correspond to our digital phalanges, to the
metacarpal, and the anterior carpal bone.
3268. The wings are the branchiæ of the Mollusc that
have been set free; they are placed therefore upon the
back and are four in number. In many Insects there is
still a pair of wing-like scales in front of the four wings,
as in some Lepidoptera. They perhaps correspond to
the shells of Molluscs, are branchial opercula.
It is only from this point of view that the structure of
Insects admits of being fully understood; and apart from
this it is absolutely devoid of all analogy. Thus it is possible
for only six legs to take their origin from the thorax
in the direction downwards, and nevertheless for there still
to be wings upon the dorsum or back. The wings of Birds
are by no means homologous to the alar appendages of
Insects; they are, as is well known, the anterior members
themselves, and there is no longer therefore in the
Bird any feet attached to the thorax below, as in Insects.
Besides, if the wings did not signify arms in the Bird,
then it must have four legs. Thus in the Insect the
wings could not also mean feet.
Their structure also speaks in favour of our view of
the Insect's wings. They are known to be completely
traversed by respiratory tubes, are true, or only desiccated,
branchiæ—aerial gills. (Ed. 1st, 1811. § 3088.)
Wings and feet are dependent from the same ring of
the body, and are thus like the branchiæ and feet of
the Crabs. Let the Crab's branchiæ elongate and dry,
and they will thus be wings.
3269. Since the wings are newly eliminated organs
of the sense of feeling, so are they here characteristic of
the animal, and are consequently of greater importance
for the purposes of division and arrangement than the
organs of the head, which in all the lower animals is
only an apparent head, and cannot therefore serve to
characterize groups, &c.
3270. That the tracheæ in Insects have been developed
out of the branchiæ by saccular inversion, is a
fact displayed in a particularly distinct manner by the
Scorpions and Spiders, who still possess at bottom internal
branchial laminæ, unto which, however, air instead
of water finds its way. It may be said, that with the general
conversion into horny texture, the arteries of the
Mussels were transmuted into internal tracheæ, and the
branchial flabellæ into external organs. Would we interpret
the body of the Insect in a strictly philosophic sense,
the parts must then receive very different names to what
they now bear. Properly speaking, in our own species
the thorax has no limbs, but only the neck. The upper
limbs are not lungs, but branchial organs, and it is the
cervical vertebræ, from between which nerves are sent off
to the arms, for it is just upon the neck also that the
branchiæ have been left. What is termed therefore the
thorax in Insects, would be properly their neck. Their
abdomen would consist accordingly of thorax and abdomen,
and it is this also, which carries on the principal
share of the respiratory process. It therefore consists
generally of ten rings, and has ten pairs of spiracula,
namely, twice five, for both thorax and abdomen.
Or again, the abdomen may be regarded as an intestinal
and sexual cavity, and the thorax may be left
with its own name. In that case there would be five
spiracles for the sex, five for the intestine, and perhaps
only two for the thorax.
Would we proceed yet further; the head can then
only be viewed as pharynx, and consequently as neck.
3271. In most Branchial animals that live in the
water, a perfect circulation is present, because, by reason
of their feeble amount of respiration, all the blood is not
consumed. This is also the case in the young of air-breathing
Insects, so long as they may have to grow.
But when they have ceased to do this, so strong an
amount of tension emerges in the circulation, owing to
the increased respiration of air, that blood remains but
in scanty quantity to be carried back by the veins, and
the arteries now for the most part convey the air in a
pure state, namely, uncombined with blood, as in the
higher animals.
3272. As the air-tubes pass to all parts of the body,
like the arteries whose place they now supply, so does
the nutritive juice become everywhere oxydized and converted
into parenchyma or tissue. The nutritive juice
doubtless transudes at once through the intestine and
penetrates to all parts, as in the Plants.
3273. Of the vascular system nothing at last remains
but the dorsal vessel, whose ramifications appear to disappear
entirely. According to its analogy with the
Crabs, Scorpions, and Spiders, it is the aorta. It appears,
as if in Insects the circulation dies off from the living
body.
The whole Insect is an air-organ, an aero-vascular
stem. All the organs respire directly, such as the intestine,
the motor fibres, nerves, sexual parts, and
wings. There is no part unto which tracheæ do not
pass, and in this respect completely resemble the arteries
of other animals.
3274. The intestine is invariably furnished with an
anus, and that too situated quite posteriorly. It is
usually expanded into several stomachs and has appendages,
almost as in Fishes, which virtually correspond to
the abdomino-salivary or pancreatic gland.
3275. The salivary ducts open into the mouth, and
thus as in the Snails and Kracken, whose similar structures
they repeat.
3276. Whether what have been called the biliary vessels
are what the name implies, and really convey bile
from the fatty body to the intestine, or whether they are
lacteal vessels and discharge their fluid near the rectum
into the dorsal vessel, does not yet admit of being determined.
They have been thought to constitute an urinary
apparatus; only certain cysts which occur in connexion with
the sexual parts, appear rather to correspond to the latter.
3277. The nervous system consists, as in the Worms,
of two ganglionic ventral chords.
3278. In the head, the feet again, and probably also
wings, undergo repetition. This repetition is nowhere
so distinct as in Insects. He who can still entertain any
doubt of the maxillæ being arms, let him resolve to
descend into the Insect-world, and he will become in
place of a sceptic, a believer. (Ed. 1st, 1811. § 3095.)
What have been called manducatory pincers move outwards
as well as the legs, and seize like arms, are only
arms. Their gripe takes place in the lateral or horizontal
direction, and resembles that of scissors or shears.
3279. Where, moreover, there are three pairs of legs
on the thorax, we find also three pairs of maxillæ,
namely, an upper and lower maxilla, and a labium or
inferior lip, which consists of the same parts as the maxillæ,
only they are united by tegument.
3280. Where there are five or more pairs of legs on
the thorax, as in the Crabs, there also are found as many
pairs of maxillæ.
3281. Upon these maxillæ, palpi are likewise situated,
which are probably nothing else than what has been
termed tarsus, thus repeated in the head also. They are
only arrested upon the upper maxillæ or mandibles.
(Ed. 1st, 1811. § 3096.)
3282. The wings appear to be repeated upon the
head as antennæ. Thus the head also is in the Insects
a perfect trunk.
3283. Upon the head there is nothing more than the
eyes. They have also become horny in texture, whereas
before they were completely membranous, as in the
Snails.
The eyes have been subordinated to the sense of feeling;
they are nervous papillæ placed beneath a transparent
tegument upon the apex of a tentaculum.
Insects therefore have a number of eyes; they stand
either separate, as in the Worms, and are then called
simple eyes, or they are crowded together, constituting
compound eyes.
3284. Of the other sensorial organs a papilliform
elevation in the pharynx is frequently exhibited as a
tongue.
3285. Ears are found in the Crabs at the root or base
of the antennæ, or in them there is only a tympanic
cavity with an ossicle inclosed within; in the Insect
there is nothing of the same kind. But meanwhile,
since many species attract each other by means of sounds,
they must thus possess the faculty of hearing. The
antennæ therefore probably correspond to the auditory
conch. The auditory conch of the Mammalia ranks also
in the signification of the hand, and thus of the organ of
feeling.
Nevertheless the antennæ may be transformations of
the wings. The auditory ossicles are indeed members
which have originated from branchiæ. Probably the antennæ
are the auditory ossicles themselves which have
emerged outwards, as in the Fishes and Reptiles.
There is no trace of a nose; nor can there be any, for it
is the anterior orifice of the vertebral canal, and this is
wanting in all lower animals. A noseless animal is an
inferior or Tegumental animal.
3286. The sexes are separate, because they are Air-breathing
animals and perfectly symmetrical in form.
The ovi-and seminal ducts are likewise symmetric and
in pairs. The ovipositors and penes are perfect, as in the
Snails, because they can be shoved forwards, but not
everted as in the latter.
3287. The position of the sexual parts at the posterior
extremity of the body has been permanently established,
with exceedingly few exceptions, as in some aquatic Insects.
That they correspond to the head, or are imperfect
cephalic organs, is most distinctly shown in the Insect.
They are usually environed by valves, pincers, and filaments,
which resemble maxillæ and palpi.
3288. The Ovum-animals multiply by division, by
granules or gemmæ, i. e. shoots, the Sexual animals by
membranous ova, the Arthritic animals by horny ova.
These last, or egg-shells, are hard, and frequently also
so strung together or laid by the parent upon each other
as to represent the annulate body of some Insect.
3289. In the ova of the Dermatozoa or Tegumental
animals the vitellus only appears to exist, being without
albumen, which first seems to make its appearance along
with the animal systems. As the animal separates into
higher and lower substances, so also does the ovum or
microzoon. The simplicity of the Tegumental animals
has been prophesied or foretold in that of their ova.
The Insecta, as being the third class of their circle,
repeat the Acalephæ and Kracken. With the last they
bear much resemblance in form and in their motor
organs.
3290. Would we arrange the Worms along with the
spiral vessels or ducts, the Crabs with the woody rings
in the stem, we must term the Tracheal animals foliage
or leaves. Their wings are pinnate leaves, and among
the Orthoptera many occur which, as well in the form of
the body as of the wings, appear as if they had been just
liberated from the Papilionaceous tree. The tales or
stories, about leaves changing in the torrid zones into
Insects, are not without meaning; for poetry is none
other than the Ideal of natural history.
Metamorphosis.
3291. We now proceed to retrace our steps. The
Insect is a Tegumental animal represented in limbs.
There can be therefore present in the Insect no other
development than in the Cutaneous animal, which works
itself up into the Branchial and Tracheal animal.
The Insect passes through three stages prior to its attaining
the adult or perfect condition. It is at first
Worm, next Crab, then a perfect, volant animal with
limbs, a Fly.
3292. The representative passage of the Insect through
the preceding classes in the course of development, constitutes
its metamorphosis. Thus the Insect's metamorphosis
obtains a meaning and an explanation. Upon the
whole, the history of every kind of pregnancy is none
other than the passage which takes place through all the
animal classes, as I have first represented in my book
upon "Generation" (Von der Zeugung, 1805); but in
no class of animals are the periods so dissevered or
drawn apart as in Insects. It therefore comes to pass
that these animals are the equivalent transcripts or
copies of a system common to them and the preceding
animals.
3293. The Worm is the first condition of the Insect.
It is represented, by the larva, which, according to its diversity
of size, is called maggot, caterpillar, grub, &c.
3294. The larva is only a Cutaneous and Intestinal
animal. It knows of nothing else but eating like the
Worm; it has no sexual function, no lust, nor pain; it
can scarcely move; in many the feet are wanting, as in
the larvæ of Flies, which thus resemble the Entozoa;
many have a crop of lateral papillæ, like the caterpillars,
which resemble the Nereids.
3295. The change into a nympha, chrysalis, or pupa,
commences with the horny induration met with in the
Crab, and in the higher organized kinds of Snail. The
pupa is the embryonic Crab or its antetype, it is the
Snail in its shell.
3296. In the third condition the Insect makes an
advance above the branchial condition, and casts aside
the Snail's or Crab's shell; and is then the perfect Insect,
the Fly.
3297. The metamorphosis is accordingly the embryonic
transition of the Insect, after extrusion from the
egg, through the three classes of its circle. In the ovum
state it only passes through that of the Intestinal and
Sexual animals.
3298. This is a retrospective proof that the higher
animals also pass in the ovum through the condition of
the lower animals, but after birth through the classes
which directly precede them.
What holds good of the Insects, does so also of the
preceding groups of animals, although in a less degree;
the higher class of each circle still passes after birth
through one or other of the inferior classes.
Thus, the Acalephæ first appear under the form of
Infusoria; next change into Polyps, and then obtain for
the first time the form of the perfect Acalephæ. The same
phenomenon occurs in the naked snails or Slugs. They
have, when freshly hatched from the egg, a small mussel-like
shell, which they lose at a later period.
The Echini or Sea-urchins, which I believe ought to
be ranged with the Kracken, also pass through a metamorphosis.
Finally, it is well known that the Crabs also are subjected
to a metamorphosis. Their first condition must be
regarded as that of the Worm.
The law is thus universal. The second and third
class of every circle traverse after birth the classes, to
whose series they belong.
It would even appear that the classes, also of a lower
circle, are subject to similar changes; at least many Entozoa,
and thus the first class of the third circle, appear
to undergo such a metamorphosis, since they first of all
resemble an Infusory animal, next change as pupæ into
Molluscs, and are then manifested for the first time as
Worms.
3299. The Dermatozoa or Sensitive animals range in
the following manner, according to the anatomical systems:
A. Intestinal animals. | B. Vascular animals. | C. Respiratory animals. |
| 1. Gastric animals. | 4. Venous animals. | 7. Reticular animals. |
| Infusoria. | Mussels. | Worms. |
| 2. Intestinal animals. | 5. Arterial animals. | 8. Branchial animals. |
| Polyps. | Snails. | Crustacea. |
| 3. Absorbent animals. | 6. Cardiac animals. | 9. Tracheal animals. |
| Acalephæ. | Kracken. | Insects. |
3300. According to the developments of the feeling-sense
they stand thus:
| A. Ovum-animals. | B. Sexual animals. | C. Cutaneous animals. |
| 1. Vitelline animals. | 4. Ovarial animals. | 7. Papillary animals. |
| Infusoria. | Mussels. | Worms. |
| 2. Albuminose animals. | 5. Orchitic animals. | 8. Pedal animals. |
| Polyps. | Snails. | Crustacea. |
| 3. Involucral animals. | 6. Renal animals. | 9. Alary animals. |
| Acalephæ. | Kracken. | Flies. |
3301. From these tables the number and kind of
relationships or affinities is readily deduced. There is a
Relationship of Proximity, as that between Infusoria,
Polyps, and Acalephæ.
A Relationship of Repetition, as of Infusoria, Mussels,
Worms. There is also a Relationship of Series, which
takes its rise from the parallelism of the classes. The
two last relationships have been confounded together
under the term analogy, which at best is but
a word of random definition.
Second Province—Sarcozoa, Cephalozoa.
FOURTH CIRCLE—SARCOSE ANIMALS.
3302. The system of motion and sensation, which
broke forth so forcibly in the tegument, now passes over
into other forms, into the globular form of bone, the
fibrous form of flesh or muscle, and the point-form of
the nerves.
3303. The nervous system when liberated from the
vegetative organs is, or consists of, the myelon (spinal
chord) and the encephalon, i. e. brain.
3304. These animals are therefore Osseous, Muscular,
and Encephalic animals, whereupon the senses are proportionately
developed, and constitute the basis of the
highest animal forms—Sarcozoa. The Dermatozoa are
asarcose or fleshless animals.
3305. With the sudden appearance of the animal
systems all organs of the head, such as tongue, nose,
ears, and eyes, are also developed; no asarcous animal
has nostrils. Now these parts, especially the nostrils, as
marking the terminal extremity of the vertebral column,
make up the head—Cephalozoa, or Cephalic animals.
Class 10. Osteozoa, Glossozoa.
3306. The animals, in which the osseous system for
the first time makes its appearance are, the Fishes.
The Fish alone has more bones than any other animal.
It has dorsal rays, which are wanting in all others.
3307. The animal systems are for the first time
slightly separated from each other. Bone, muscle, and
nerve are rather a gelatinous mass, which only bids fair
for becoming something higher; on this account the
bones are frequently but soft cartilages or tendons, the
muscle composed of white fibres like those upon the
intestine or vessels, the nerve thick, oleaginous, and soft
in texture, while the brain is in its constituent parts
hardly comparable with that of the Thricozoa.
3308. The muscles of Fishes are not yet perfect in
character, since they are devoid of individualization and
red colour, and their fibres run mostly parallel with each
other without uniting into tendons. Their muscular
body is a muscular wall.
3309. The Fishes, ranking upon the first stage of the
Sarcozoa, repeat the same stage of the preceding circle,
and thus the Infusoria, Mussels, and Worms; or the
stomach, veins, and branchial rete or network; furthermore,
the vitellus, ovary, and skin, which systems must in
them accordingly predominate.
Pelvis.
3310. In Fishes the pelvic organs, sexual parts, and
tail predominate. The tail, as being an appurtenance of
the pelvis, is larger and stronger than in other animals.
It mostly makes up the largest part of the body, and
is, properly speaking, its only motor organ.
3311. The sexual parts have still the form of the
intestine, and occupy the greatest part of the abdomen.
The ovaria are two sacs, like two Infusoria, in whose
parietes granules are developed; even the testes are
only two such sacs or seminal vesicles wherein the "milt"
is contained. Fishes may be termed anorchitic, or animals
devoid of testes.
The ova are small, consisting of spawn or roe without
shell, but they separate into albumen and vitellus.
3312. External sexual parts are not present. Everything
usually opens into a cloaca, which is thus here a
true pharyngeal cavity.
3313. With the perfected formation of the head, the
animal, so to speak, undergoes a sudden and entire change,
and the sexual parts are developed with all their accessory
organs, such as the sexual lung. Kidneys are present
and mostly an urinary cyst. The kidneys are indeed
still so amorphous and soft, that they more resemble
coagulated blood than an organ, yet meanwhile agree in
this respect with the 'milt' and branchial substance.
Abdomen.
The abdomen is not simply confluent with the sexual
body, but thrust by it completely forwards. The anus
mostly lies in front of the middle of the body.
3314. The intestine separates for the first time distinctly
into cephalic and sexual, or small and large,
intestine.
3315. The vitelline canal is the cœcum. This is very
distinct in the Sharks and Rays.
3316. In Birds therefore the vitelline canal is the
cœcum also, and the two cœca, falsely so called, which
are placed upon the side of the rectum cannot represent
the cœcum coli, or else the Bird must have three of these
blind appendages or sacs.
3317. The spleen here appears for the first time;
while the pancreas divides into a number of cœcal appendages.
Thorax.
3318. The thorax of the Fish is reduced within very
narrow limits; it is like the first thoracic formation, and
is thus a Mussel's thorax. The branchiæ with their
opercula are similarly formed to the branchial plates
and shells of the Bivalve Mollusca. The thorax is therefore
attached only externally to the body, and the Fish is
to be viewed as a Mussel, from between whose shells a
monstrous abdomen has grown out.
3319. But this Molluscan thorax is conjoined with
animal systems, and has assumed their noble or elevated
type of structure. Here therefore the osseous and sarcose
system blend together, and the higher formation of
the thorax emerges into view.
3320. For the first time an advance is made towards
the formation of a trachea, namely, by the branchial
framework or skeleton, which opens into the mouth,
and corresponds therefore properly to the larynx. Fishes
are therefore the first animals which breathe through the
mouth. In all the preceding classes the air entered the
body, or the water found access to the branchiæ, by other
routes.
3321. They may be termed Mouth-breathing animals,
for the first formation of the trachea extends no further
than to its junction or communication with the mouth;
since, for it to be continued into the head and open self-substantially
as a nasal organ, constitutes a second step
in advance, which cannot be ventured upon in an Abdominal
animal. In Fishes everything relates to the
abdomen, and this is expressed by the first union of the
trachea having taken place with the pharynx or the
mouth.
3322. But, at the same time that the trachea is inserted
within the mouth, it opens externally and in the
lateral direction upon the body, thus letting the water
escape from it posteriorly after a mussel-like or sexual
fashion. Thus, the trachea is not yet closed inferiorly,
nor hence also the thoracic cavity. Between the head and
abdomen there are still openings—branchial foramina.
3323. The trachea is itself, however, a thorax upon a
small scale, consisting as it does of rings, or as it were
ribs. These arches are not yet united by muscles with
each other, and the water flows out between them into
the apparent thoracic cavity, from which it next escapes
beneath the branchial operculum.
3324. These tracheal rings are the branchial arches.
The branchial vessels are tracheal and by no means
pulmonary vessels.
3325. Thus, upon taking a retrospective glance at the
matter, we may fairly conclude that the branchiæ of the
Dermatozoa are not equivalent to the lungs of the higher
animals, but are only the antetypes of the bronchi, being
thus cervical organs.
3326. The trachea is thus formed prior or antecedently
to the lungs, but is still membranous and devoid
of any continuous connexion between its rings.
3327. The lung is an organ extraneous or foreign to
the trachea, and becomes only as if accidentally associated
with it.
3328. But a Sarcozoon is not devoid of lungs nor of
aerial respiration; for, being the totality of all lower
animals, it consequently unites in itself the respiratory
apparatus of the Branchial and Tracheal animals; thus
gills and lungs.
3329. In the Fish the first lung appears, that is to
say, if we designate or interpret the respiratory organ in
Insects only as tracheæ, which do not open into the
mouth.
3330. The Fish's lung is the air-or swim-cyst.
3331. The lung is still subordinate to the abdomen
and intestine. It is therefore still separated from the
trachea or the branchial arches.
3332. Fishes swallow the air and drive it into the
pulmonic cyst, where it is analysed or decomposed.
3333. This lung, so soon as it stands opposite to and
enters into conflict with a branchia, is what serves to
direct or control the action of the heart. It is only when
the swim-bladder is allowed to hold good as lung,
that the circulation in Fishes admits of being understood.
For otherwise venous blood must flow into the heart, from
this into the branchiæ, from these directly into the aorta,
and so to the different organs without again entering a
cardiac cavity, a structure which occurs in no other class
of animals, but rather in every instance its reverse. The
first heart is arteriose, not venous in all animals, even in
the Mussels and Snails and in the embryos. Now, the
Fish is still such an embryo, and has only an arteriose
heart.
3334. Thus, the following is the "modus operandi"
of the parts in question; the swim-bladder is the
lung, in which, seeing that it contains air, the blood is
oxydized; this oxydized blood flows into the heart and
renders it arteriose in character, despite the concomitant
influx of venous blood into that organ. Upon this, the
blood passes out of the heart through a true aorta, which
is called the branchial artery. Instead then of this aorta
giving off only some twigs as bronchial vessels to the
gills, and next proceeding to pursue its course as a main
trunk through the body and along the back, it passes
wholly to the branchiæ, i. e. it entirely becomes a bronchial
vessel, is slightly oxydized, and then returns upon
itself to form the aorta, which should have taken its
course directly from the heart.
Senses.
3335. In accordance with the thorax, the corium or
true skin is developed. It is mucous in character and
muco-secernent like the intestine, from its being constantly
in the water. The whole skin is undermined by
mucous canals and perforated by their excretory apertures.
These foramina of the lateral line are arrested,
and metamorphosed branchial foramina, which have only
retained the evaporative function of respiration.
3336. The remnants of the tegument's annulation
are the scales. They are desiccated air-branchiæ,
alary opercula or elytra, and consequently indicate that
feature of the Insect, which has continued to work its way
into the class of Fishes.
3337. Like the skin, so are the limbs—tegumental
members. What is flesh and bone upon them, has kept
quite close to the body, and that only which is designed
for splitting into digits projects, constituting tegumental
digits with cartilages—fins. These fins are something
better than the lateral papillæ of the Worms, are
furnished at their base with joints, and are in number
only four, but crippled or stunted in every possible way.
3338. The fin-rays do not correspond to the digits,
but to the nails. They are fibrillated nails, like the
wing-feathers of Birds.
3339. Lastly, the head possesses all the organs of
sense which belong to a head, but they are still far
removed from their perfect condition.
3340. As the nervous system is the first mass, from
which the remaining ones have been set free, so also is
the nervous sense the first after that of the tegument,
which appears as a whole, and serves as a pattern for the
subsequent senses. The eye is the sense which is first
developed most perfectly, not directly as regards its own
completeness, but when considered in reference to the
others.
As the sense of feeling has been at once manifested in
the vegetative animals as a peripheric nervous sense; so
antithetically or vice versâ it is in the higher animals the
light-sense, as being the central nervose sense, which
appears.
3341. Hitherto both these senses stood altogether
alone with each other upon the stage of animal existence,
in order, as it were, to play their parts in mutual consent;
but so soon as the sense of feeling became individualized
into tactile organs, then also the eye made its
appearance in a detached or isolated manner.
3342. The Fish's eye is upon the whole composed of
the same structures as in the Mammalia; but it is devoid
of motion and external coverings.
3343. The ear, as being the sense of motion, has
scarcely withdrawn itself from the brain, has not yet
become a true external organ, and that portion of it
which appears externally, is subservient to inferior systems,
such as the branchiæ or gills.
3344. The external auditory organ has become confluent
with the branchial aperture, and the auditory
ossicles have become parts of the branchial operculum.
3345. In the internal ear only the three semicircular
canals have been left. The cochlea is not yet developed.
As true palpebræ or eyelids are wanting to the visual
organ, so also are the conchs of the external ear.
3346. The nostrils exist, because a vertebral canal,
which terminates in them, is present; stoutly-developed
olfactory nerves are also present, so that the sense of
smell cannot be wanting. Only this sense has not yet taken
the respiratory organ into communication with itself,
and both therefore live for themselves in an arrested or
stunted condition. This nasal organ does not open
into the mouth, admits of neither water nor air passing
through it, and therefore serves not as a test-organ to the
respiratory process. This is my main point for distinction
of Fishes from Reptiles.
3347. Every Sarcozoon, whose nostrils do not open
into the mouth, is a Fish. The Siren therefore does not
belong to the present class, while the Lepidosiren, having
an imperforate nasal organ, does.
3348. The tongue has remained rather an organ of
touch and deglutition, than an instrument of taste. The
salivary glands are scarcely developed.
3349. But the tongue here appears for the first time
in the animal kingdom as a perfect organ provided with
muscles, and a lingual bone, as in Man; Fishes are, as
regards the development of this sense the Glossozoa,
Lingual or Tongue-animals.
3350. As Fishes are the repetition of the intestine and
vitellus, so they may be termed abdominal or rather
Pelvic animals. They are an abdomen, unto which are
appended branchiæ, fins and head.
They are the repetition of the Infusoria, Mussels, and
Worms; in them we meet with mucus, branchial opercula,
articulated filaments, and labial cirrhi or palpal
organs.
Class 11. Myozoa, Rhinozoa.
3351. Those Sarcozoa which for the first time obtain
true muscles and a perforated nasal organ are the
Reptilia.
3352. True muscles are red, have a definitely circumscribed
outline, and are divisible into head, fleshy portion
or belly, and tail or tendon. As such they are found for
the first time in Reptiles.
Thorax.
3353. With the osseous system in Fishes the sexual
abdomen was chiefly developed; with the muscular
system therefore, as in the present class, the abdomen
proper or intestinal abdomen becomes perfectly developed,
and the thorax more spacious in calibre.
3354. The thorax is still blended with the abdomen.
In Fishes the disposition was already stirring to create for
themselves an air-organ; only this was imperfectly
attained, since the swim-bladder communicated with
the gullet, but not with the branchial arches or larynx,
nor with the nasal organ, as in the present class.
3355. Now, if the swim-bladder be symmetrically
developed, if it communicates with the branchial
larynx and opens through the nose, the aerial respiration
is completely attained, and shares its dominion over the
body along with the digestive function.
3356. A Fish, respiring through the nose into two
swim-bladders, is an Amphibious animal, a Reptile.
3357. In the Reptile, however, the thorax is still
subordinated to the abdomen. Its lungs traverse the
whole abdominal cavity, and its manner of breathing is
still similar in character to the abdominal function. The
lungs are simply two membranous bladders, like two
intestines, and admit also of being voluntarily filled with
air like the intestine is with water. This filling or inflation
of the Reptile's lungs takes place also by the abdomen
being expanded at the same time that the mouth is
closed by the muscles of the fauces, and is therefore a
true process of deglutition.
3358. Their inspiration is still therefore fish-like in
character, excepting that the air is drawn in by the
nostrils; the respiratory process, however, and the respiratory
organ is constituted as in the perfect air-breathing
animals.
3359. Reptiles may therefore be called Abdominal
animals, while the Fishes are Pelvic animals.
3360. The metamorphosis of the branchial arches into
larynx is placed beyond doubt in the Reptiles. The anterior
branchial arches frequently unite with the lingual
bone, whereby the latter obtains several cornua.
3361. The thyroid gland also makes its appearance
here for the first time, while the branchial vessels separate
from the arches. Fishes have therefore no thyroid
gland.
3362. The circulation is complete. The venous and
also the arterial blood enters the heart. But both kinds
of blood are still mingled with each other as in Fishes.
Yet already, through the direction and arrangement of the
cardiac orifices, provision seems to be made for their
separation.
3363. The rationale of this mixture of the blood appears
to reside in the fact, that many of these animals,
and probably all of them in the ovum state, breathe by
branchiæ. (This proposition, which was announced in
the 1st Edition, 1810. § 305, and there based upon the
transitional stages traversed by the animal classes during
embryonic development, has since then been raised to a
state of certainty.)
3364. The mixture of the blood takes place through
an opening in the septum separating the two cardiac
ventricles, and which corresponds to the foramen ovale
of the fœtus. The heart of Reptiles is therefore a persistent
fœtal heart.
3365. Without doubt, however, the arterial blood
only enters the left heart, while the venous remains in
the right, to be thence driven into the lungs.
3366. In other respects the foramen ovale is not a
true opening, but a bifurcation of the vena cava, one
branch of which passes into the right, the other into the
left ventricle, as I have demonstrated in the 'Beyträgen
zur Anatomie' (Frankfurt bei Wesche), in treating of
the calf's heart.
3367. There are generally no foramina in the body,
but only fissures without orifices.
Abdomen.
3368. The digestive organs are pretty similar to those
in Fishes, as is exemplified by the intestine, liver, and
spleen; the digestive process is, however, more energetic,
and frequently aided by venom or poison.
3369. The sexual parts are perfect. The first true
testes and ovaria have originated. The tegumental formation
has consequently passed over at the extremities
into a glandular, because the vascular system preponderates.
Even in the highest Fishes, such as the Rays and
Sharks, the testes and ovaria are still not developed to the
extent which they are in Reptiles.
3370. In the female the oviducts pass together as
complete cornua into an uterus. In the male the seminal
ducts are still indeed separate, yet the two bodies or
crura of the penis are frequently developed, and continue
separate in the Serpents and Lizards, but are connate in
the Tortoises.
3371. The kidneys are distinct and symmetrical; their
ureters usually unite into a large urinary cyst. Thus,
the sexual lung is also significantly developed.
3372. Many of these urinary cysts split, as in the
Tortoises, into two cornua. This is the form under
which the bladder again occurs in Birds, where its two
cornua have been incorrectly termed cœca, but the urinary
bladder rectum, because the intestine opens into it.
3373. In Reptiles also there is a cloaca or sexual
orifice, into which all the sexual apertures meet together.
The ova of Reptiles possess a vitellus and albumen,
and are surrounded by a membranous or sometimes by
a calcareous shell.
3374. Hence Reptiles have originally also a cœcum,
but it has for the most part disappeared; like its antetype
the vitelline canal, which is so small in most Fishes and
Birds, as to be scarcely distinguishable.
The tail projects from the body, and is rather trailed
after the animal, than enabled to assist the body in its
motions, as in Fishes.
Senses.
3375. Many Fishes have only a vertebral column without
lateral bones; in the Thoracic animals this deficiency
can hardly occur any longer, although we are not, on the
other hand, to expect any great amount of perfection to
be attained by these parts. Reptiles have osseous
ramules, whether such be ribs or true feet; the Serpents
have indeed only ribs, but these exist in great numbers;
in the Frogs, on the contrary, ribs are wanting, but they
have limbs.
In no Reptile do fin-rays occur; consequently, there
are no peculiar bones as in Fishes.
3376. In Reptiles the limbs themselves are in a varied
or undetermined state, although they constitute a class
in which the formation of true muscular limbs is decided.
In Reptiles the toes appear for the first time.
3377. As regards the tegument, it quite corresponds
with that of the Corals, Snails, and Crabs, of which the
Reptiles are a repetition. In some cases it is slimy and
naked; in others Mussel-shells originate, as in the
carapace of the Tortoises; the scales, scutes, and claws
of Serpents and Lizards repeat the rings and scutes of
Crabs.
3378. The head has separated itself more from the
thorax, the eyes and ears being much more perfect; as
regards the latter organs the cranium is always perforated,
in the former there are eyelids.
3379. The eyes have still no power of free motion, and
the eyelids are as yet very imperfect. It is principally
the lower eyelid which closes, while in Mammalia the
upper one is the most developed.
3380. The external auditory meatus is notwithstanding
usually covered by integument, the auditory ossicles
are mostly blended into one, which projects above the
skull; the cochlea is wanting.
3381. Reptiles produce sounds for the first time
through the lungs; they have voice, but as yet no song.
3382. The nose, however, gives the finishing touch to
the head. It has not only opened as a vertebral canal
in front, but also as a thoracic cavity posteriorly into the
mouth, which was not the case in Fishes. The open
nose is the æsthetic character of Reptiles, just as the
red muscles are the characteristics of the anatomical
systems. The Reptiles are Rhinozoa.
3383. The tongue takes a higher rank than in Fishes.
It is throughout soft, fleshy, and smooth; but in most
species is still slit into two, which reminds us of a similar
condition of the penis.
3384. The teeth are in these animals more like digits
than in Fishes. While here they consist for the greatest
part of front teeth, and are therefore associated with the
intestine, in Reptiles they are mostly lateral, and thus
true maxillary teeth, which are annexed to the salivary
system. With this special dental formation the saliva is
also more active; it is a rapidly fatal poison.
3385. The poison-teeth or fangs have likewise a groove,
which can be regarded as the continuation of the salivary
duct.
Class 12. Neurozoa, Otozoa.
3386. Those Sarcozoa, whose nervous system is for
the first time perfectly developed, and whose ear is open
externally, are the Birds.
3387. The completion of the nervous system is the
brain; now the brain defines or determines the head;
the Bird is, properly speaking, the first Encephalic
animal.
3388. In the Bird the head has for the first time, and
that indeed suddenly, freed itself from the trunk, and
been placed upon a long neck, far removed from the
thorax. In no class are there such long necks and so
many cervical vertebræ to be found as in Birds. They
can be therefore called also Cervical animals, as the neck
is not simply present, in order to render the head independent
or self-substantial.
The caudal vertebræ, on the contrary, are lessened
throughout the Birds, to a degree met with in no other
class of animals.
3389. The brain separates for the first time completely
into cerebrum and cerebellum, begins to exhibit
convolutions, and has in general most of the individual
parts analogous to those in the Thricozoa. They are
therefore Encephalic animals, which the name of Neurozoa
will properly enough imply. The nerves are in
comparison with the spinal cord much thinner than in
Reptiles and Fishes.
In the Bird also, all the spiritual or mental faculties
make their appearance for the first time and suddenly,
whereas in the preceding classes but slight traces of
them were observed. Such for example are their mechanical
instincts, varied modes of nidification, powers of
imitation, susceptibility to instruction, knowledge of
their benefactors, sentiment of joy, wheedling or coaxing
manners, and so on. We have no example of Fishes and
Reptiles having learnt any artificial tricks.
3390. Birds are the closest repetition of the Insects,
but stand remote from the Crabs and Acalephæ, indications
of which are reflected by their structure, disposition,
mechanical instincts, and in their nests.
Thorax.
As the sexual body with the tail predominates in
Fishes, and the abdomen in Reptiles, so does the thorax
in Birds. The whole abdomen and sexual body has
been subordinated to the thorax.
The ribs are here directed in such a manner for the
first time, that the thorax can act as a voluntary pump-organ.
Sternum and respiratory muscles are usually
large and peculiarly built.
3391. The lung is only a cluster of Insect tracheæ.
They are full of foramina, out of which the air can penetrate
into the whole body, exactly as in Insects. In Fishes
the lung was still an actual intestine, as seen in the single
air-bladder; in Reptiles it was nothing more than a
double intestine; in Birds this lung is divided after the
fashion of an Insect into aerial vessels or ducts.
In the Bird the intestine lies in the air, and breathes
from it, as in the Insect. Birds are also animals which
breathe by the intestine.
3392. The whole Bird is lung. Its body is a thoracic
cavity, while in the Fish it was simply a sexual cavity,
and in the Reptile had obtained the abdominal form.
In Birds therefore there is a number of ribs and a
strong ossification, from the air itself penetrating into
the bones.
Even the intestine has passed over into a motor organ.
In Birds and Insects only do we find a true muscular
stomach, wherein the food is crushed.
Members or Limbs.
3393. In these animals the formation of limbs or
members must be wholly attained. Everything, that
works, must work to the production of limbs. The whole
body becomes limb.
3394. With the perfected nervous and respiratory
system the bones and muscles make also a perfect appearance.
The skeleton is hard, complete, and full of air
instead of marrow; the muscles are red and separated,
the movements free and complex.
3395. The elevated condition of the motor system is
demonstrated in the limbs, which here appear in the
greatest diversity. There are only two pairs of limbs,
thoracic, and abdominal or sexual members. These two
are equivalent, so long only as the sexual and abdominal
cavity belonging to the trunk proper predominate, and
therefore preserve the equilibrium. But the limbs, which
have been given to serve the office of the trunk, are destined
for progression or swimming, and are simply terrestrial
or aquatic members.
3396. The higher limbs are thoracic respiratory members,
which are filled with air and clothed with tracheæ,
thus exercising a function conformable to the thorax.
The thorax has an aerial character. The highest thoracic
limbs must be aerial members.
3397. As the abdominal members move upon the
earth or in the water, so do the thoracic in the air. The
terrestrial limbs are feet, the aerial limbs, wings.
3398. The wings are in the member-formation the
extremes of the thoracic limbs. It does not follow from
this that they are the noblest in rank, but only that they
are the uttermost unto which bodily motion can attain.
3399. The winged animal is the Bird.
3400. The Bird is an Insect with fleshy limbs.
3401. The wings of Birds repeat the alary appendages
of Insects in the flesh.
3402. A Bird's wing is a strange but very instructive
composition. It consists, namely, of a Reptile's foot and
an Insect's wings.
3403. We saw how the branchiæ of the Insect dried
up, separated from the feet, and being liberated as wings,
were permeated by tracheæ. In the Bird these wings
have remained standing upon the feet and been converted
into feathers.
3404. A feather is an Insect's wing.
3405. As the Bird grows out at the thoracic limbs
into the Insect's wings, so does it upon the whole body
into dried branchial laminæ.
The whole body of the Bird is clothed with branchial
laminæ or plates.
3406. The wings of the Insects might be called free
tracheæ.
The Bird's feathers are Insect-tracheæ. As in the
Insect the wings are a leash or net of tracheæ held
together by membranes, so feathers are tracheæ dividing
into fibre-like ramules.
3407. The Bird is a Reptilian or Frog's body, beset all
over with Insects, like as with parasitic animals.
3408. The highest Insect only attains to the possession
of four wings, which in some Moths split again into
several feathers. In the Bird a vast number of such
wings originates.
3409. An Insect's wing is not more than one feather,
and is therefore placed also directly upon the body.
These wings must multiply so soon as they occur upon a
membered trunk, upon arms. Thus, we need not ask
why the Butterfly has four, but the Bird only two wings,
seeing that the latter is the nobler animal. The discourse
cannot be concerning the wings of the former,
for the Butterfly has indeed none, but only four
feathers.
3410. What the Bird is, it is by virtue of its feathers.
It is throughout a trachea, a pair of bellows. Its bones
are hollow, full of air, and stand likewise in communication
with the lungs; the feather-quills are also hollow.
Senses.
3411. The wings have all the muscles to themselves;
the bone has in them gone to ruin. On the legs, on the
contrary, the muscles have declined, and the bone has
got the upper hand.
Hence it results, properly speaking, that only the thoracic
members are perfected, because the Bird is nothing
but thorax. The abdomen has, so to speak, vanished,
and through this the abdominal limbs are left only, as
thin and dry poles or staffs.
3412. From the same cause the muscular flesh upon
the head has disappeared. Neck and head are lean, or
like Insect horn, which serves only the nervous system.
3413. Beyond the excess of motion the sense of feeling
has been nearly lost. The toes are simply destined for
motion, and to be used as scrapers, and the digits have
become the supports of feathers.
3414. The bill is an Insect's proboscis. In the Bird
no teeth whatever project from the flesh, but the jaws
themselves. To such an extent has the flesh been withdrawn.
What has been called the cere is the only remnant
of the facial flesh. Even nostrils and tongue have
suffered ossification.
3415. The tongue is a feather. Saliva is scarcely
present.
3416. The ears, as being the sense of motion, are far
more perfectly evolved than in all the preceding classes.
They have opened outwards, and possess an additional
auditory part, the cochlea.
With the limbs the sense of hearing must of necessity
grow perfect.
3417. The Bird is thoroughly, or "out-and-out"
organized as an animal of song. In it Nature attains
unto a definite hearing and speech. The Bird speaketh
the language of Nature.
With the Bird, the voice, properly speaking, breaks
forth for the first time, and that too in a high grade of
perfection, as melody.
3418. The ear is the highest representation of the
trachea in muscles and bones. The Bird is the Otozoon.
Sexual Parts.
3419. The kidneys are symmetrically constructed,
although not yet a perfectly coherent mass. They are of
very large size. But a strange feature has come to pass
in the urinary cyst, which is the sexual lung. Into
this, as has been already said, the intestine opens, and
thus it here again passes over into the lung, just as
it has passed over into a fleshy cardioid stomach, and
completely upwards into the feather-like tongue.
Into this sexual lung the seminal ducts also enter, or
the penis, when one is present, along with the oviduct.
3420. The ovum consists of two completely separated
substances, and these indeed are so distinct, that the
vitellus is secreted at an entirely different place to the
albumen, the former in the ovarium, the latter in the
oviduct—whereas in Fishes both originated together.
The separation cannot extend further, or else the substances
would no longer unite with each other, and the
vitellus must be first mixed with the albumen, after it has
been completely formed, after or when it is an embryo.
In the Bird albumen and vitellus come together during
their passage out of the parent's body, or in the act of
being laid, and thus before the albumen has been converted
into the chick. But in the Mammalia they are
first completely united, subsequently to their being laid—i. e.
during the period of lactation.
3421. The vitellus is directly secreted from the arteries,
but the albumen from an enteroidal sac or the
oviduct, which is finally converted into mammary glands.
3422. The vitellus is more a product of the thorax,
and is therefore formed directly into the intestine and
entire embryo.
3423. The albumen is an intestinal or digestive product,
a solution of the organic mass into protoplasma
or primary mucus. It is not fashioned itself into the
embryo, but is only absorbed as fluid nutriment by the
latter.
3424. The shell of the ovum is the last bone, which
the animal deposits from the sexual blood, as being analogous
to the urine. It is an aerial product, or an analogue
of the feather's quill.
3425. Even the nest of the Bird is a spiritual repetition
of its plumage, for in it the stalks of plants, tracheæ, or
feathers are united into one body, which reminds us in
the Swallows of the Acalephæ.
FIFTH CIRCLE. AISTHESEOZOA.
Class 13. Aistheseozoa.
3426. An animal with all its organs of sense perfectly
developed, is a Thricozoon.
3427. The nervous system emerges at length freely
above the other systems, and it is no longer its mass, but
its organs which impart character to the animal. The
nervous organs are, however, simply the organs of sense.
Through these therefore must the present differ from
the preceding class of animals.
3428. Now too the senses first make their appearance
in a self-substantial manner above the other organs,
serving merely their own functions, and only by chance
those of others.
3429. As in Birds the whole body was subordinated
to the thorax, in Reptiles to the abdomen, in Fishes to
the sex, in Insects to the tracheæ, and so on; in like
manner is it here subordinated to the system of the
senses or the head.
3430. As the higher senses determine or define the
anterior part of the head, and are in their state of perfection
provided with muscles; so here the face or visage
is invested by flesh, whereby, properly speaking, a true,
namely, a moveable countenance, first originates. The
Aistheseozoa have a fleshy face.
3431. All possess moveable eyes; fleshy noses, which
stand open both externally and internally; ears opening
outwards, and mostly provided with a moveable flap; a
fleshy tongue, free in front, and moveable lips; with at
least thoracic limbs and a skin covered with hairs.
3432. In the Bird, Reptile, and Fish the face is
merely invested by tegument, nearly devoid of any
muscles, and therefore immoveable. They have a tegumental
face, which is incapable of producing any
expressions.
3433. In the tegumental face the eyes are motionless,
and very rarely both directed so forwards that they can
view an object together; the nostrils are frequently
devoid of a fleshy rim; the tongue often feather-like,
cartilaginous, or covered with teeth; true fleshy lips are
wanting, as are frequently the teeth, with even the limbs
and digits, or these are divided into a number of rays,
forming either feathers, or fins; in the Aistheseozoa there
are never more than five toes present, and if there be
fewer, then the number admits of being referred to some
crippling of the normal quantity, five.
3434. It is remarkable, and serves to the discovery of
many laws, that the highest sense here appears for the first
time in its state of perfection. The eye in the Aistheseozoa
is present in a perfect condition throughout the class,
with the exception of the eyelids; the other organs of
sense are, on the contrary, exhibited in all their gradations
of structure.
3435. It appears, as if the whole animal was first
perfected, when the eye is present with all its investment
or clothing. The eye of the Aistheseozoa has not simply
all its internal chambers and all its humours, but also
all its muscles; it is moveable and has perfect eyelids,
with very few exceptions—Ophthalmozoa.
3436. The ear now begins to suffer arrest. Its completion
is indeed the formation of an external concha or
flap for receiving the rays of sound, the hand being
repeated in the ear, and its skeleton constituted by the
auditory ossicles. This auditory hand occurs only in the
Aistheseozoa, and might serve as characteristic if it were
not wanting in many, while the eyelids are present. As
in the Whales, where the auditory passage can be, however,
closed, an act which is not possible in any Bird.
The Bird must hear whether it will or no.
In all Thricozoa the interior of the ear is perfect,
having cochlea, semicircular canals, tympanum, and, as
brachial parts, the three conjoined auditory ossicles.
The concha of the ear passes besides through all stages
of development, from having a simple margin to one with
most varied convolutions, lobes and opercula.
3437. Still more than the ear does the nose undergo
modifications. In the Whales it seems to be less adapted
for the purposes of smell than of respiration. The olfactory
nerves are in them very delicate, and the apex of
the nose is destitute of motion.
In other beasts, on the contrary, it is elongated into a
very muscular proboscis or snout, and is endowed with
the power of voluntary motion.
The form also of the nostrils is very varied, they being
round, narrow, patulous, and frequently capable of being
closed.
3438. The tongue is indeed mostly fleshy and soft;
yet in many species it is provided with horny points, in
others invested by a dense coriaceous tegument, so that
it appears to represent an instrument for deglutition
rather than gustation.
3439. The lips also are mostly fleshy and moveable;
yet in many they retrograde exceedingly and lose their
mobility, as is partly the case in the Ornithorynchus.
3440. In most, however, the limbs, especially the toes,
are still subject to variation. Their perfection consists
in the number five, and in the difference between the
two pairs of limbs, as in Man. In the Apes the posterior
feet are also hands, which is an imperfection; in the
Marsupials there are hands posteriorly but toes in front;
in other respects there are generally toes, sometimes five,
then four, finally two perfect and two dew-claws in the
Ox, and at length only one in the Horse, while the posterior
extremities are virtually lost in the Whales.
3441. The dental system, as being the set of maxillary
claws, is alone present in its state of perfection in
the Thricozoa. They alone have, in addition to the
incisor teeth, all the five kinds of teeth that differ from
each other in form, namely, canine, false molars, laniary,
with second and third true molars, corresponding to the
five fingers reckoned from the thumb.
3442. In the dentition of those animals which tear
their food the greatest amount of completeness and variety
is met with, since each tooth has a different form and
function.
In the Bears the molars are uniform in character, and
so on through the Apes up to Man.
In the Marsupials they are tolerably similar, as also in
the Bats and Shrew-mice.
They are still more alike in the Pig and Horse, and the
incisor teeth begin to be wanting in the Ruminantia,
e. g. the Ox.
In Mice the canines are wanting, in the Sloths the
front teeth, and in the Ant-eaters actually all.
3443. In opposition to the perfect eye the general
sense of feeling is developed in the tegument. The
tegument which is best developed will be that, which
represents a self-substantial organ with all its appurtenances,
and thus an organ of touch, whose nobility of
rank consists in motion. A skin, which is moveable by
means of muscles, must take the noblest rank. A skin
with tegumental muscles is an organ of feeling, which is
already in some degree subjected to the influence of the
will. If tegumental muscles do not occur in all these
animals, they still do in most of them.
The production of the most perfect coat or covering is
the second step, whereby the tegument ascends.
3444. Hitherto the outer covering of animals was
pretty inorganic, consisting either of hollow tracheæ,
feathers, or semicavernous scales, coat of mail, or lastly,
only mucus. All these organs were only constructed
after the type of individual vegetable systems, in greatest
part only after the respiratory organ, and were therefore
partial in aspect or deficient.
The highest covering must also bear the highest
meaning. This is that, which grows out of the sanguinary
system collectively, out of the capillary vessels.
The capillary vessels of the covering are, however, the
hairs. The hair is the most perfect covering of the
animal.
3445. These animals are the Thricozoa.
Already the highest birds, e. g. the Ostrich and Cassowary,
exhibit feathers which merge over into hairs.
3446. An integument clothed with hair, and self-substantially
moveable is the perfection of this organ, it
is a fur—Furred animals.
3447. The fur is the peripheric combination of the
Vegetable and Animal. The hairs are the highest Vegetable,
as being the vascular system grown out of or above
the animal, and constituting the fundamental or basi-system
of the whole body. The tegumental muscles are
the lowest Animal. In the fur consequently the whole
animal has been represented, but as a limit set between
animal and world.
3448. The tegumental covering also varies. The
hairs become scanty in respect of number; instead of
them spines, horny rings or scales occur, yet there are
always hairs on the belly. In many Whales they seem
to dwindle into bristles. In some they have coalesced to
form a kind of bark, as in the Stellerian Walrus or Sea-cow.
Thorax.
3449. In all the respiration is aerial; in all there is a
true costal structure, and a respiration effected through
the motion of the ribs or a pumping action; in all the
lungs are filled with cells; in all there is a diaphragm,
a larynx, a trachea with cartilaginous rings, and a thyroid
gland. But the air no longer penetrates from out the
lungs into all the cavities of the body, as in Birds.
Abdomen.
3450. The two intestines, namely, the large and
small, are more distinct from each other than in other
animals; the cæcum coli is in most distinct; the stomach
expanded and membranous in texture, so that it operates
simply through chemical influence. Liver, pancreas, and
spleen occur in all.
3451. The sexual parts are in every respect very perfect.
The penis is present in all, and all have an uterus,
fallopian tubes, and separate oviducts. The penis is in
many still retracted into a cloaca, and the testes frequently
lie within the abdominal cavity.
3452. As in the tegument the parts separate and
each becomes self-substantial, the fibre animal, the tegument
vegetal, and so on even in the sexual animal. The
ovarium, consisting of two parts, now also separates, since
one becomes animal, but the other remains vegetal. The
albumen-organ becomes animal, detaches itself from the
sexual parts, is developed in the self-substantial tegument
and called mamma.
The Aistheseozoa are thus also Mastozoa or Mammalia.
3453. This separation of the sexual animal is one of
the first characters. The mammæ could never be wanting,
because they indicate an essential stage in the
development. The albumen-organ becomes an organ of
sensation. If the existence of mammary organs be
doubtful, as in the Ornithorynchus, the hairs are in that
case perfectly distinct. No Reptile nor Bird can have
hairs, because their covering is derived only from a
partial system, the respiratory; while hairs are from the
general vascular system, which is the foundation of the
tegument, of the sense of feeling. The Ornithorynchus
is a furred animal, and this would suffice to bring it
among the Aistheseozoa, were we even to deny it mammary
organs.
3454. Both sexes, male and female, have breasts.
In youth they are most readily detected in the male,
because their sexual parts then rank nearer to the female.
In other respects the mammæ are probably the chiefest
organs of absorption for the embryo.
3455. The mammæ are nobler in rank, when, as
udders they become self-substantial; and the more they
remove from the belly or abdomen, and come, as breasts,
to be placed upon the chest.
D. ORDERS AND FAMILIES.
3456. Had the names been given in strict accordance
with rank; then the class-divisions, which correspond to
the circles, would have been called Orders.
3457. Families would be class-divisions, which correspond
to the classes themselves.
3458. In no class therefore could there be more than
four orders admitted, or five, if the sensorial organs be
reckoned to constitute a special circle.
From the same reason there are no more than thirteen
or seventeen families.—These names cannot meanwhile be
so strictly adopted, because the classes are not of equal
rank, as has been observed; on this account it is necessary
to shove in here and there other divisions, which
should be termed at one time Cohorts, at another, Alliances.
3459. If the animals of a class differ from each other,
it is only possible by their bringing to bear in addition
to their characteristic organ some other organ, and consequently
becoming similar to an earlier or later class.
3460. Yet nevertheless in this ascent the animals
could never outstep the confines of their own circle.
There can be no Dermatozoon which could have bones;
for in that case it would be an Osteozoon, and belong to
another province.
Every class therefore comprises as many orders only as
there are circles with which it comes into contact. Thus
in the first circle there is only one order, in the second
two, and so on.
Each class therefore includes also as many families
only as the circles which are touched by it contain classes.
Thus the first to the third class has three, the fourth to
the sixth, six; the seventh to the ninth, nine; the tenth
and so on, thirteen.
3461. The serial arrangement of animals into families
is naturally difficult; but in Physio-philosophy we have
to treat not about the execution of the Systematic in
detail, but concerning its principles.
FIRST CIRCLE. INTESTINAL, OOZOA—MUCUS-ANIMALS.
First Class.
Gastric, Vitelline Animals—Infusoria.
3462. The Infusoria admit of being reduced to three
divisions: the lowest of which, such as the Infusoria proper
or Monades, are provided for the most part with cilia;
the next in succession, as the Rhizopoda, possess extensible
processes, and are mostly covered by a multi-chambered
shell; lastly, the most perfect Infusoria have all kinds of
internal organs, and especially what has been called rotatory
apparatus, as being the dawn of future tentacula.
3463. These three Families obviously correspond to
the three classes of the present circle, and that indeed as
follows:
To the first class, or Infusoria proper, correspond the
Monades.
To the second class, or Polyps, the Rhizopoda.
To the third class, or Acalephæ, the Rotifera.
3464. The Monades are obviously the simplest organized
creatures, being mucous vesicles, which move, obtain
their food by stirring up vortical currents in the water,
and emit what is undigested again by the mouth.
3465. They occur very abundantly in all infusions,
and can very well originate, like Fungi, by division of the
organic mass, although they are in a condition to propagate
themselves, i. e. by spontaneous division.
3466. The Monades are the semen of the animal
kingdom, which is dissolved in, or rather produced from,
the sea.
3467. The animal body is nothing else than a compound
fabric of Monads.
3468. Decomposition is a separation into Monads, a
retrogression into the primary mass of the animal
kingdom.
3469. All propagation, even that of the sex, commences
like the animal kingdom, or with its first family.
On that account the embryonic development must be a
passage through the animal kingdom.
3470. The Rhizopoda usually adhere together within a
multi-chambered calcareous shell, out of which they protrude
mucous filaments, and are therefore the antetypes
of the Corals or Polyps.
3471. The Rotifera exhibit all kinds of viscera, such
as intestine and ovaries; besides what have been called
rotatory organs, which remind us of the arms or tentacles
of the Acalephæ.
3472. The families of these animals may be therefore
aptly named as follows:
Fam. 1. Typical Infusoria—Monades; cilia or vibratile organs.
2. Polypary Infusoria—Rhizopoda; extensible processes.
3. Acalephan Infusoria—Rotifera; intestine and oral organs.
Second Class.
Intestinal, Albuminous Animals—Polypi.
3473. The Polyps also do not admit of being divided
into more than three families. The first are only tubes
or vesicles with capillary tentacula around the mouth, as
the naked Polyps, Tubulariæ, Sertulariæ, and Cellulariæ.
The others have true ciliated tentacular rays surrounding
the mouth, and are always condensed inferiorly
into a horny, and occasionally stony axis or stem, as is
seen in the Gorgoniæ, Alcyoniæ, and Isidiæ.
The third have ordinary tentacula, occurring in great
numbers like tassels or tufts around the mouth; their
integument is either hardened into stone, or becomes
fleshy, as in the Star-corals and Actiniæ.
3474. There is no doubt that the naked Polyps are
closely allied to the Infusoria, and among them indeed to
the Rotifera, so that they represent but a higher stage of
these creatures, characterized by superior size and by
tentacular in place of ciliary or vibratile hairs.
3475. The Cellulariæ cannot be more distinctly characterized
than by saying that they are cortices or ramules
inhabited by Vorticellæ. Thus they are Vorticellæ surrounded
by a shell, and may be compared to ova, in
whose coriaceous or leathery shell calcareous granules are
blended, as in the eggs of the Crocodiles and Tortoises.
3476. They increase by means of ova and by ramification,
if the former mode of division is not carried too far.
3477. The tubes of the Tubulariæ appear to be
nothing else than the posterior extremity of the Polyp,
desiccated or dried. These tubes are not therefore the
product of an excretion, but the body itself.
3478. On the contrary, the tubes of the Sertulariæ
must be held as being a tegumental excretion, within
which the Polyp ramifies and produces ova-cysts. If the
naked Polyps resemble ova which are devoid of shells,
such as the roe or spawn of Fishes, then the Sertulariæ
resemble ova surrounded by a tough skin, like those of
Rays and Serpents.
The Radiated Polyps or Horn-corals are invariably
ramified and converted towards their interior into a common
horny or stony mass; so that the animals themselves
appear to have coalesced upon this into a common tegument
or bark. They possess a stomach and ovaries surrounding
it, which open into the border of the mouth
between the rays. They thus increase by ova and ramifications.
They therefore represent the class proper of the
Polyps.
The tufted Polyps comprise the proper Corals or
Lithozoa, are in form and substance like the Acalephæ
but with this difference, that the covering is mostly
lithoidal, while in many Acalephans, as the Porpitæ, it
only appears as a cartilaginous disc or plate.
3479. These Corals are true ova, having a perfect
calcareous shell, like that of the Bird. The gelatinous
animal which adheres within a wide-mouthed Madrepore,
e. g. Fungia, resembles a vitellus just hatched, and from
which the fœtal envelopes have been developed.
3480. The numerous tentacular filaments surrounding
the wide mouth resemble the shags of the chorion, which
accumulate around the orifice of the umbilical cord to
constitute a placenta in the higher animals.
The Corals are brood-eggs inclosed within the uterus
of Nature or in the sea. The Coral-animals are an umbilical
cord, treasured up for the embryo, while the
Tubulariæ are only membranous vitelli, and the Gorgoniæ,
ova with desiccated albumen.
3481. The families of Polyps stand therefore in the
following order of significance:
Fam. 1. Infusorial Polyps—Tubulariæ.
2. Typical Polyps—Alcyoniæ.
3. Acalephan Polyps—Actiniæ.
Third Class.
Absorbent, Involucral Animals—Acalephæ.
3482. The Acalephæ also can only be brought under
three divisions, viz. Röhrenquallen, or Physaliæ, Rippenquallen
or Beroes, Hutquallen or Acalephæ.
3483. The Physaliæ are without doubt the lowest,
being only giant Infusoria; the Beroes are allied through
their simple bodily cavity, to the Gorgoniæ; the Acalephæ,
through their form, to the Madrepores, especially the
Actiniæ.
3484. They range therefore in the following order of
significance:
Fam. 1. Infusorial Acalephæ—Physaliæ.
2. Polypary Acalephæ—Beroes.
3. Typical Acalephæ—Acalephæ.
3485. The first have not yet attained to the unity or
single character of the mouth, but imbibe their food
through numerous tubes. They are bundles of ramified
Vorticellæ, a thoracic duct full of glands and roots, which
absorbs nutriment out of the sea, instead of an intestine.
As the first family they are the antetypes of the Mussels
and Entozoa, and especially of the Hydatids and
Tape-worms, and we shall not be far from hitting the mark,
if we compare their air-bladder with the hindermost cystiform
member of the body in the Hydatids.
3486. The second family have a single mouth and
ribs mostly full of lamellæ, which are probably the antetypes
of gills; they are also frequently traversed by nutritive
vessels.
They are antetypical of Snails, being in form an
abdominal pouch, in substance a liver, and frequently
possessing paired tentacula. Higher up, and they indicate
the Crabs.
3487. The third have, as a general rule, one central
mouth surrounded by four arms and very numerous vessels,
which run from the gastric cavity to its margins,
and there mostly elongate into filaments or hairs, being
thus true lymphatic vessels, which convey the nourishment
directly from the stomach through the whole body,
and become on its margin tentacula.
3488. As antetypes of the Kracken they particularly
manifest the form of the Sepiæ, or Cephalopods, in the
strong and frequently papillated arms that surround the
mouth. Gland-like nodes are also developed in the
margin of the pileus, but their meaning is as yet doubtful.
Upon a higher stage, and they are antetypical of Insects.
Their vascular system forms an exceedingly regular,
quaternary net, with branches and ramules ranging
opposite to each other, which have been believed to be
the vitelline network of the ovum when hatched. These
vessels terminate likewise in a common marginal vessel,
as in the vitellus.
3489. Viewed as a whole they resemble in form, appendages,
and substance the fœtal involucra or envelopes.
The upper surface may be compared with the convex
back of the envelopes, the lower to the concave funnel
of the umbilical cord, and the succigerent filaments or
hairs to the shags of the chorion. They are probably
elongated by injection, like the so-called legs of the Star-fishes.
The secretion of the ova in four ovaries speaks also in
favour of their higher grade of development. They lie
in four cavities around the stomach, which also open
near to the mouth. Their size also is significantly expressive
of their higher position.
Finally, they are of separate sexes. In them is the
milt for the first time distinctly displayed, and in the
same situation indeed where in other animals the
ovaries are.
3490. Hence, in these three classes there are found
no more than three families, which together make up
only one order. The want of a fourth family and so on,
proves that the development of these animals does not
pass over into the succeeding circle. Their body therefore
is only an homogeneous, transparent mass, variously
excavated and perforated, but not separated into two
cysts, namely, tegument and intestine, and without the
other viscera, which are formed by the vascular system,
such as the liver, kidneys, and salivary glands.
SECOND CIRCLE. VASCULAR, SEXUAL ANIMALS.
3491. These animals will both traverse the three
classes of their circle, as also repeat the preceding three
classes, and they consequently divide into two orders
and six families.
Fourth Class.
Venous, Ovarial Animals—Mussels.
3492. The Mussels or bivalve Mollusca resolve themselves
into two orders, according to the structure of their
mantle or respiratory sac. It is either closed in a
tubuliform manner, and opens posteriorly into two
mostly tubular-shaped respiratory apertures, and in front
has an orifice for the passage of the foot—Camacea
(Lochmuscheln); or it is slit along its whole length
anteriorly; and the pedal aperture is confluent with the
anterior and also indeed with the posterior respiratory
opening—Ostracea (Spaltmuscheln.)
The first order still represent the cystic form of the
Oozoa, and partly by tubuliform calcareous shells placed
around the two ordinary shell-valves; but the second,
by the freer development of the organs, represent the
animals of their own circle.
Order 1. Protozooid Mussels—Lochmuscheln.
3493. These animals repeat the Mucus-animals; are
tubular in form, and mostly provided with two long
respiratory tubes.
Fam. 1. Infusorial Mussels—Sackmuscheln or Pholades.
Cylindrical, with a tubuliform mantle almost entirely
closed and frequently surrounded by a calcareous tube,
in addition to the two ordinary shell-valves; the pedal
aperture is situated at the oral extremity; the respiratory
tubes retractile by means of muscles; the foot is cylindriform.
Of the same kind are the Pfahlmuscheln, Teredines,
and Sandmuscheln, as also the Solenidæ. They typify
the Nudibranchiate Snails and the Salpæ.
Fam. 2. Polypary Mussels—Klaffmuscheln or Tellinidæ.
Body flat, mostly discoidal; there is a large fissure
on the abdominal side of the mantle for the passage of
the foot, while posteriorly there are two long respiratory
tubes with retractor muscles. Here belong the Tellinidæ
and Venusidæ.
I have shown, that they are already to be recognized
by the uncinate or hook-shaped groove for the mantle
in the shell, and which proceeds from the insertion of
the retractor muscles. The foot is usually lancet-shaped.
They typify the Patellæ and Ascidiæ.
Fam. 3. Acalephan Mussels—Cardiacea.
Body globular in form, mantle having a pedal slit on
the abdominal side; two respiratory apertures without
retractor muscles; foot mostly uncinate or strap-shaped.
T have shown that these creatures are to be recognized
by a discoid groove in the shell, which simply depends
upon the mantle, the retractor muscles being wanting.
Here belong the Cardiacea and gigantic Chamidæ, in
which last a byssus occurs, and the union also of the two
occlusor or sphincter muscles, as in the following order.
They antetypify the air-breathing Snails and Cirripedia.
Order 2. Conchozooid Mussels—Spaltmuscheln.
3494. These animals represent their own circle.
The pedal slit in the mantle occupies the whole length
of the latter, and is confluent with the anterior or also
with the posterior respiratory aperture; so that only one
or even no aperture is left, and this when present is
besides never elongated in a tubular form; the retractor
muscles are therefore wanting also, and the shell simply
exhibits a discoid groove for the mantle. The occlusor
muscles approximate and unite at their terminations.
Both usually coalesce into one.
Fam. 4. Typical Mussels—Mytilaceæ.
Occlusor muscles separate, and mantle slit to such an
extent, that only the posterior respiratory aperture remains;
foot mostly tongue-shaped or coniform, with occasionally
a byssus.
Here belong the fresh-water Mussels and the Mytili
proper, of which last many bore into rocks. They
typify the Capulidæ and Brachiopoda.
Fam. 5. Snail-Mussels—Arcaceæ.
Have two separate occlusor muscles and a perfectly
separate mantle without respiratory aperture; the foot
small, mostly cartilaginous. Here belong the Arcæ,
Aviculæ, and Pearl-mussels.
They typify the Trochidæ and Pteropoda.
Fam. 6. Kracken-Mussels—Ostracea.
Only one occlusor muscle, mantle entirely open or slit,
so that both respiratory apertures are only oblique incisions
therein; foot very small, frequently furnished with
a byssus. They typify the Buccinidæ and Cuttle-fish.
Fifth Class.
Arteriose, Orchitic Animals—Snails.
3495. The Snails likewise divide into two orders,
according to the two circles, preindicated by the Ovum-and
Sexual system. Their branchiæ are either ramulate
or pectiniform, the sexual parts combined or separated.
The first kind are still frequently gelatinous in texture,
transparent and naked; their branchiæ usually stand out
freely as filaments, lamellæ, or ramules upon the back, or
lie simply as a vascular network within the mantle. All
are androgynous. They therefore obviously repeat the
Protozoa or Mucus-animals.
The second are invariably covered by the shell and by
a mantle, within the cavity of which the branchiæ lie concealed
as one or two comb-like bodies. Tentacula and
eyes, which are occasionally wanting in the preceding
order, are here universally present; the sexes separate.
As in the Snails, the male parts make their appearance
for the first time distinct and individualized, and are also
a characteristic organ of the class; so also do they serve
as bases of division, and the Snails may be divided into
those which are Androgynous or bisexual, and those
with separate sexes, i. e. Diœcious.
Order 1. Protozooid Snails—Androgyni.
3496. The male and female sexual parts united together
in a single individual, branchiæ ramuliform, occurring
either as filaments, leaflets or ramules, freely situate upon
the naked body, or as a network lodged within the pallial
cavity and surrounded by a shell.
The Nudibranchiate species live in the sea, those with
hollow branchiæ or the Pulmonea in the air; the former
feeding mostly upon animals, the latter upon plants.
Fam. 1. Infusorial Snails—Notobranchiata, or Tritoniæ.
Body gelatinous and membranous, cylindrical and
naked, devoid of shell, with branchial filaments or ramules
disposed in two rows upon the back.
Here belong the Tritoniæ and Dorides. Their body
is muscular; the tentacula not retractile; the male
sexual parts open in company with the female upon
the right side of the neck, as in the higher organized
Snails. All inhabit the sea. They prefigure or typify
the Salpæ.
Fam. 2. Polypary Snails—Pleurobranchiata, or Patellæ.
Body and sexual parts as in the preceding family,
the branchiæ, however, occurring as ramules or leaflets
upon the sides of the body, are more or less covered.
Here belong the Phyllidiæ, Schüssel-and Schildschnecken.
They are the antetypes of the Ascidiæ.
Fam. 3. Acalephan Snails—Dictyobranchiata or Limacidæ.
The branchiæ form a rete or network within the
pallial cavity, and respire the moisture of air; mantle
and viscera are mostly surrounded by a shell; the body
is therefore bipartite, being separated into a splanchnic
or visceral body and a foot with head.
Here belong the Air-breathing Snails, both land as
well as fresh-water species. They typify the Cirripedia.
The shells are mostly thin and horny, yet nevertheless
contain a considerable quantity of calcareous earth, and
are mostly devoid of opercula.
Those species, which dwell in fresh water, do not
possess introvertible tentacula like the marine Snails,
and the eyes are placed at their basis; the sexual orifices
are separate.
In the Land-snails, the tentacula are introvertible, and
support the eyes upon their apex; the sexual apertures
are blended into one. The former, like the marine
Snails, lay numerous small ova inclosed within a gelatinous
mass in the water; the latter deposit their ova free,
covered with a membranous, and occasionally calcareous
shell, in the earth. Copulation is effected in all by a
reciprocal interchange of the androgynous species.
I have observed that individuals of Limnæa auricularia
can continue to propagate for several generations
without being impregnated; they must impregnate themselves.
Order 2. Conchozooid Snails—Diœcii.
3497. Branchiæ situated within the cavity of the
mantle, and hanging down in the form of a comb; shell
for the most part spirally contorted; sexes separate—Pectinibranchiata.
Here belong the Capulidæ, Turbinidæ, Neritæ, Conidæ,
Volutidæ, Buccinidæ, Muricidæ and Strombusidæ.
The tentacula are not retractile, and have the eyes
mostly seated at their basis; the penis is external, very
large, and cannot be drawn within the body, but is only
reflected into the pallial cavity; most of them have a
protrusile perforating proboscis, and an operculum. They
lay numerous small ova, contained within large membranous
cases, which frequently hang in rows to each
other like a necklace of pearls. The shells are in some
instances horny, in others of a stony texture.
Fam. 4. Mussel-Snails, Capulidæ.
Only one branchial comb within the mantle, and
covered merely by a flat hood-shaped shell; no operculum.
Here belong the Capulidæ; all the species are marine.
They prefigurate the Brachiopoda.
Fam. 5. Typical Snails, Turbinidæ.
Two branchial combs, mantle devoid of a respiratory
groove, shell turbinated; mostly furnished with an operculum,
and that indeed of a stony texture. Here belong
the Turbinidæ, and Trochidæ, such as the Cyclostomata,
Paludinæ, Ampullariæ, Janthinæ, Neritæ. Live in the
sea and in fresh-water.
Fam. 6. Kracken-Snails, Buccinidæ.
Like the preceding family, but their mantle has a
groove, and there is a horny operculum. Here belong
the Conidæ and Volutidæ, Buccinidæ, Muricidæ, Strombusidæ.
Almost all live in the sea and are of sanguisugal
habits. They antetypify the Cuttle-fish.
Sixth Class.
Cardiac, Nephritic Animals—Kracken.
3498. Body cylindrical, without sole or foot; frequently
venous hearts together with the arteriose hearts,
and a kidney-like organ.
Such are the proper Cylindrical Snails, which are
either sessile or fixed, or move themselves by fins and
what have been called arms—Sessile and Natant Kracken.
Here belong the Salpæ, Cirripedia, Brachiopoda, Pteropoda,
Heteropoda, and Cephalopoda. They all live in
the sea.
Branchiæ and sexual relations very varied.
They divide likewise into two orders, in accordance
with the two circles of Mucus-animals and Conchozoa;
the former are spathiform, mostly gelatinous, and firmly
sessile or fixed; androgynous, without head and rudders
or fins. The branchiæ trellis-like and filiform—Ascidiæ,
Cirripedia and Brachiopoda.
The others have a kind of head with arms and eyes,
or fin-shaped steerage-organs upon the body; branchiæ
pectiniform, reticular, and leaf-shaped—Pteropoda, Heteropoda,
and Cephalopoda.
Order 1. Protozooid Kracken—Rumpfkracken.
3499. Body sacciform, mostly gelatinous, without
head and eyes; species androgynous. They are destitute
of independent locomotion, most of them being quite
stationary or fixed, and surrounded frequently by a shell;
some few float about passively in the sea.
Fam. 1. Infusorial Kracken—Salpæ.
The Salpæ are gelatinous, freely natant cylinders,
perforated by an open tube, within which are situated
the branchiæ, heart, mouth, intestine, and liver, without
any tentacula. They continue to hang for a long time
to each other, as though they were in the ovary.
They undergo a remarkable metamorphosis, which
traverses two generations, the young meanwhile not
resembling the mother but the grandmother.
Body within a sacciform mantle, with two opposite
respiratory apertures.
Fam. 2. Polypary Kracken—Ascidiæ.
The Ascidiæ have a sacciform body with two co-approximate
respiratory apertures; branchiæ internal and
trellis-shaped.
These gelatinous or cartilaginous animals adhere
firmly to rocks, with the respiratory apertures directed
upwards as in the Mussels, but the branchiæ are not
foliiform; the internal cavity is lined with a membranous
sac, upon which the branchial vessels are dispersed in a
trellis-like manner. At the bottom of and within the sac
is the mouth, which is destitute of tentacula. Intestine,
liver, and a heart that is simple or undivided. Mode of
propagation unknown.
They are frequently connato-sessile, like the Polyps.
Fam. 3. Acalephan Kracken—Cirripedia.
Body sacciform, without branchial openings, oral apertures
situated at the inferior extremity of the longitudinal
fissure in the mantle, and provided with a kind of
maxillary apparatus; above or behind the mouth are
annulate or horny filaments representing caudal fins or
feet, upon the basal joints of which is a branchial filament.
Here belong the Balani and Lepades—Fuss-kracken.
These animals are likewise sessile, at least when full
grown, and are surrounded by shell-plates, which remind
us of the Echini or Sea-Urchins, whose antetypes they
are; this resemblance is particularly distinct in the
Balani, which have similar odontoid valves upon the
aperture or mouth of the shell. These animals also
undergo a metamorphosis, since in the early periods of
their existence they swim about like small Crustacea,
and later on become fixed or sedentary, and surrounded
by the plates of shell.
The Lepades have the ordinary shells of Mollusca, but
each of the two valves consists of two pieces, between
which an odd one is interposed down the back.
These five shell-pieces become in the Balani a regular
set of teeth, which are adherent within the cylindrical
shell, by which the posterior part of the body, or the
peduncular and elongated mantle, is surrounded. (Vid.
Oken's 'Allgemeine Naturgeschichte,' V, 1835, 509.)
They bear a resemblance to the Crustacea, not simply
in the annulate horny feet, but also in the double and
ganglionic nervous chord lying along the abdomen; only
they are androgynous, which occurs in no Crustacean.
Order 2. Conchozooid Kracken—Kopfkracken.
Tentacula upon the head or fins upon the body. They
repeat the Conchozoa or Shell-animals.
Fam. 4. Mussel-Kracken—Brachiopoda.
The body is surrounded by a mantle open above, and
by two shells; on the sides of the mouth there are two
arms.
These animals look exactly like Bivalve Mollusca;
they are firmly adherent to rocks, being frequently supported
upon a hollow pedicle, which is a prolongation of
the mantle. Branchiæ and sexual parts but little known;
species probably androgynous—Armkracken.
3500. In the following families the body is free and
sacciform, provided with arms or fins, and mostly with
eyes upon a kind of head.
These animals have the mouth situated above or in
front of the body, and are distinctly separated into
mantle and abdomen, while they are frequently surrounded
by a shell. They have a kind of head with
tentacles or arms, and frequently furnished with eyes
like the Snails, or pretty nearly like the Fishes; the
sexual parts united and separated. They all row, paddle,
or steer themselves about in the sea.
Fam. 5. Snail-Kracken—Pteropoda.
Body mostly gelatinous, sacciform, closed all round
and free; fins upon the sides of the neck, with two tentacula
frequently projecting from near the mouth; androgynous,
naked, and inclosed within a shell.
These transparent animalcules swim about upright in
the sea, and wave the fins in this position as rapidly as a
butterfly does its wings. Most of them are covered with
a spathiform and likewise transparent shell. The branchiæ
are placed externally on the body, but are not very
distinct. Several species have an appendage in front of
the neck, which is obviously a rudiment of the Snail's
pad or sole—Flugelkracken.
The Heteropoda appear to resemble the Plantar or
Sole-Snails, but the body is mostly gelatinous, and the
foot or sole is only compressed to form a fin, so that they
can only swim but not creep with it. Many of them
have a shell almost like that of the Argonauta. Pterotracheæ—Ruderkracken.
Fam. 6. Typical Kracken, Cephalopoda or Sepiæ.
Muscular animals inclosed within a sacciform mantle,
open in front; head furnished with eyes, and surrounded
by more than four strong prehensile arms; laminated or
leaf-like branchiæ within the mantle, sexes separate; in the
body is a nephroidal or kidney-like gland, which secretes
a dark-coloured or inky fluid, on which account they are
called by us 'Dintenschnecken'—Sternkracken. They are
obviously the highest organized Malacozoa, and already
remind us strongly of Fishes, partly from their size,
partly by their fleshy body, and in part by their perfect
organs of vision.
The body is often as large as the trunk of a man; the
head being separated from it by means of a neck, has in
its interior a kind of cartilaginous brain-case or skull,
with externally a pair of jaws resembling a Bird's beak,
and with eyes tolerably similar to those of Fishes. Auditory
organs are also present, which consist of a tympanic
cavity inclosing an otolithe or ear-ossicle; nostrils are
wanting. The so-called arms are perfect organs of locomotion,
as also useful for seizing the prey, being provided for
this purpose with sucking cups, which adhere by producing
a vacuum, when applied. In other respects these arms are
nothing else but the Snail's sole divided in front into
lobes. The ova resemble berries, and hang attached to
one another in branched clusters like a bunch of grapes.
The Sepiæ possess a remarkable gland, which is complicated
with the liver, and secretes a dark brown juice,
which has been called ink or sepia; it probably ranks in
the significance of the kidneys.
Many are covered with a shell externally, as the
Nautili and Argonautæ; but in the common Sepiæ this
shell adheres within the mantle upon the back, and
forms a straight lamina or plate, which is sometimes
horny, sometimes calcareous in its texture. It has been
called the "white fish-bone." What is regarded as the
dorsal side of the animal is its ventral side, because
upon the former lies the pallial aperture and the anus,
while the vitelline sac is also inserted there.
In the form of the body, as well as in the circular
position of the cephalic arms, the Sepiæ strikingly
resemble their antetypes, the Acalephæ and Cirripedia,
and their metatypes, the Asteriadæ and Crustacea.
With these animals the second stage or circle through
which the Dermatozoa pass, or that of the Sexual animals,
is closed. They require only a slight additional grade of
perfection, and they would pass over into another class
of animals. Thus were the arms to become horny and
articulate, they would be Crustacea; had the head a
nose, and the body consequently a myelon or spinal chord,
they would then be Fishes.
SECOND CIRCLE. RESPIRATORY, CUTANEOUS ANIMALS—ANCYLIOZOA.
3501. Body annulate.
Here range the Worms, Crustacea, and the proper
or volant Insects.
3502. In this circle a remarkable relation is revealed.
If, namely, its classes, orders, and families be compared
with those of the two preceding circles, it is then distinctly
shown that the present one again commences quite
from below, ascends parallel with both these, and transcends
them in its highest inmates or members.
As an example of the first case, the imperfect condition
of some Entozoa, and their great resemblance to Infusoria,
is sufficiently expressive.
The Worms evidently pass parallel to the Mucus-animals,
and the Entozoa indeed to the Infusoria, the
Red-blooded Worms to the Polyps, but the Holothuriæ
chiefly to the Acalephæ, near which they are still arranged
up to the present day, although they are articulated, and
have an intestine and vessels.
Thus this class traverses the three inferior classes, and
consequently imparts to them only the value of orders, or
must itself be raised only to the rank of an order, a step,
however, which, considering the great number of Worms,
would not be suitable or just. The names may therefore
remain for the sake of uniformity; only it must be borne
in mind that they are of unequal value.
The same relation is exhibited in the Crustacea or
Crabs. They obviously admit of being parallelized with
no other animals than the Malaco-or Conchozoa. The
Entomostraca and Crabs evidently repeat the Mussels,
the Aselli the Snails; but the Spiders and Scorpions the
Kracken or Cephalopoda. Consequently here also the
orders correspond to the preceding classes.
The true or proper Insects transcend the Sexual animals,
and therefore conform to the classification of their own
circle, namely, they repeat the Worms and Crabs, and
finally mount forth upon their own stage.
The inferior or lower organized animals form consequently
two series, viz. the Smooth and Annulate, which
in their lower members range parallel, and that indeed
after the following manner:
| A. Smooth. | | B. Annulated or Ringed. |
| I. Protozoa. | | I. Worms. |
| a. Stomach, | 1. Infusoria. | | 1. White-blooded Worms. |
| b. Intestine, | 2. Polypi. | | 2. Red-blooded Worms. |
| c. Absorbents, | 3. Acalephæ. | | 3. Holothuriæ. |
| II. Conchozoa. | | II. Crustacea. |
| d. Veins, | 4. Mussels. | | 4. Crabs. |
| e. Arteries, | 5. Snails. | | 5. Aselli. |
| f. Hearts, | 6. Kracken. | | 6. Spiders. |
| III. Ancyliozoa. | | III. Flies. |
| g. Retia, | 7. Worms. | | 7. Tracheoptera. |
| h. Branchiæ, | 8. Crabs. | | 8. Dictyoptera. |
| i. Tracheæ, | 9. Flies. | | 9. Ceratoptera. |
The Worms have simply an annulate body with reticular
or filamentary branchiæ, but no feet; the branchiæ
are supported by the body.
The Crabs have an annulate body with feet; the
branchiæ are supported by limbs.
The Insects have an annulate body with feet and
wings; the body is supported by the branchiæ.
Seventh Class.
Reticular, Papillose Animals—Worms.
Protozooid Ancyliozoa.
3503. There are Worms with a soft body and white
blood, without proper tentacula; others with red blood,
with and without tentacula, having also bristles along the
sides of the body; lastly, the rings of the body become
hard or muscular, and a circle or ring of osseous maxillæ
forms around the mouth, which is usually surrounded
by numerous tentacula, as in the Star-fishes and Holothuriæ,
in whom the blood is nevertheless colourless.
They divide accordingly into three orders, and each of
these again into three families, in accordance with the
classes, namely, those of the Protozoa or Mucus-animals,
through which they pass.
Order 1. Infusorial Worms—Parasites.
3504. The White-blooded Worms or Entozoa are very
imperfect animals, a fact which proves that this circle
recommences quite inferiorly or from below. They live
for the most part in the interior of other animals, being
thus located in darkness and in places where they obtain
but little oxygen. Their blood therefore is not simply
colourless, but even its vessels are only imperfectly
developed. They respire without doubt through the
tegument. In many, a distinct or separate intestine
being wanting, it is the tegument also which digests; in
others the intestine is simply a sac without an anus. The
sexual parts too are in many species of a doubtful character,
while meantime there are both androgynous
and diœcious individuals. In the latter the male parts
open invariably at the posterior extremity of the body, as
in Insects; the female parts in front of its caudal end, as
in the Crustacea. Both are in other respects constructed
as in Insects, namely, there are two oviducts or seminal
ducts, which unite before reaching their external orifice.
They divide into three groups.
In one of these the body is tolerably smooth, and the
pharynx commencing from a simple suctorial mouth elongates
into a ramified intestine without an anus; they are
androgynous—Saugwürmer.
In others the body itself supplies the place of the stomach;
it is corrugated, and receives the food through
one or several orifices, without separating into a special
intestine. They have nearly all a claviform proboscis or
rostrum, with which they perforate substantial cavities for
themselves; it would seem that they are both androgynous
and of separate sexes—Hydatids and Tæniæ.
Others are separated into tegument and intestine like
the Mussels and Snails, but without having a distinct
vascular system, heart, and liver, though provided with a
nervous chord and separate sexual parts—Ascarides.
Fam. 1. Monad-like Worms, Saugwürmer.
Body tolerably smooth, suctorial mouth and sucker,
the intestine losing itself in the tissue of the body, and
without an anus; androgynous.
They remind us through their small size and structure
of the Monades, especially the Cercariæ, and among the
Worms typify or prefigurate in particular the Hirudines
or Leeches, both by their form and power of suction, as
also in the ramification of the intestine.
Many Cercariæ might be metamorphosed into Distomata,
and so be or constitute their young.
The Planariæ are slightly different from the Liver-flukes
or Fasciolæ hepaticæ, since they are white, bloodless,
have a ramified intestine without anus, and increase
also by fissiparous generation. They belong to the present
order, although they live not as parasites, but in a
state of freedom.
Fam. 2. Rhizopodoid Worms, Hydatids.
Body annulate or articulate, with a claviform proboscis
without an intestine; they appear to have several
mouths.
The Cystica develop ova simply upon the internal wall
of the tegument; the body's last ring is expanded into
a cystic form, and is mostly much larger than the whole
body. The Cœnurus cerebralis, or Brain-hydatid, ramifies
like the Polyps.
The Cestoidea have in each joint or member of the
body an ovarium, and, as it would appear, male parts
also, so that they are androgynous. The sexual orifices
are at the border or upon the surface of the posterior
members. The member can be viewed as an ovarium
liberating itself; as in the Lerneæ and Arguli.
The Filariæ or Thread-worms, which also belong here,
are in the form of a cylindrical tube, with separate sexual
parts, which open posteriorly.
Fam. 3. Rotiferal Worms, Ascarides.
Body cylindrical, intestine free, with mouth and anus;
sexual parts separate.
Upon the mouth these Entozoa have some papillæ or
points, which may be regarded as tentacula. The larger
species have a distinct nervous cord. The Gordius cannot
well take any other position than the present, although it
does live in a state of freedom.
Order 2. Polypary Worms—Hirudines.
3505. Body cylindrical with branchiæ in or upon the
tegument, blood red; two ganglionic nervous threads
run along the ventral surface of the body, as in Insects;
all are indeed androgynous.
Here belong the Red-blooded Worms, as Hirudines,
Lumbrici, Nereides, and Serpulæ.
Fam. 4. Tubularial Worms—Dermatobranchiata.
Body cylindrical. The branchiæ are only a vascular
network in the tegument.
Here belong the Hirudines and Lumbrici.
The Hirudines have a perfectly naked body, without
filaments and bristles, a perfect network of vessels containing
red blood within the tegument, an intestine with
an anus, both sexual parts androgynoid; posteriorly
they have a pad or sucker; in the mouth there are mostly
maxillæ, and upon the head simple eyes.
The Naiades and Lumbrici have bristles in longitudinal
rows upon the sides of the body. The latter are
androgynous. The Naiades increase by division.
The Thalassemæ have a protuberant white body; red
blood-vessels only upon the intestine; mouth rostriform.
These Worms stick in the mud, and are nourished by
it. They have here and there bristles, but which do not
form any longitudinal rows. They appear to absorb
water through the skin, and respire by the intestine.
They cannot, by reason of their red blood, range with
the Holothuriæ.
Fam. 5. Alcyonioid Worms—Notobranchiata.
Body depressed, with pedal filaments, tentacula and
branchiæ in two longitudinal rows upon the back.
These Worms all live in the sea, adhering mostly
upright in the earth, and have frequently annulate,
rigid tentacula, ocelli, and pairs of maxillæ, like Insects.
The gills are not unfrequently covered with pergamentaceous
scales, as in many Nereides, and especially in
Aphrodite.
Fam. 6. Actinioid Worms—Auchenobranchiata.
Body inclosed within a pergamentaceous or calcareous
tube with lateral bristles, branchiæ and tentacula upon
the neck or head.
The neck is surrounded by a kind of mantle, almost
as in the Snails, so that several animals have been
arranged here, which are now known to be veritable
Snails. Upon the head many have a horny operculum,
whereby they can close the shell.
Here belong the Amphitritæ, Terebellæ, Serpulæ, and
Sabellæ.
Order 3. Acalephan Worms—Sternwürmer.
3506. These creatures attain the most perfect structure
of the Worms. The blood is white, the form cylindrical,
globular and stellate; the mouth surrounded by a
ring or wreath of maxillæ. The nerves form a ring
around the pharynx, and upon the latter are placed
membranous cysts, which spirt water into the tentacula
or feet as they have been called, and thereby expand
them.
It is impossible for these animals to continue to range
with the Acalephæ, although they resemble them in
outward form; for they consist of two cysts, the intestine
having become freed from the outer one as a special sac;
moreover, they have a perfect vascular system, distinct
muscles, a mouth with a dental apparatus, which prefigurates
a complete skeleton, a peculiar vascular system
for the injection of the tentacula or feet, a nervous ring
surrounding the pharynx, an ovarium entirely separated,
and lastly, a perfectly annulate body.
Fam 7. Physalial Worms, Holothuriæ.
Body cylindrical, very plentifully supplied with muscles.
Mouth and anus present, the former of these being surrounded
by a dental wreath and ramified branchimorphous
tentacula; branchiæ upon the intestine, and branchial
aperture at the anal extremity. They correspond to the
Physaliæ.
Their tegument is a perfect muscular tunic, consisting
of several longitudinal bands, which pursue a downward
course upon its internal surface; in other respects being
transversely corrugated and full of papillæ, partly replete
with hollow podoidal filaments, which by the injection
of water into them are elongated; thus, they present a
resemblance to the Acalephæ.
Fam. 8. Beroeal Worms, Echini.
Are at bottom Holothuriæ with ossified tegument, or
Asteriadæ with shortened rays.
The Echini have an anus, pretty like that of the
Holothuriæ, and also similar feet, which are protruded
in a similar manner through apertures in the shell.
The dental wreath surrounding the mouth is a complete
bony framework, which has been called, on account of
its form, "Laterna Aristotelis," and bears a considerable
resemblance to the valves of the Balani, the antetypes
in general of the Echini, just as these are of the Opossum
Shrimps.
Fam. 9. Medusal Worms, Asteriadæ.
Body stelliform, consisting of bony rings; contains a
free multilobular intestine with blood-vessels and branchiæ;
several ovaria; it would appear that they are without
male parts.
Here belong the Encrinites, Pentacrinites, and Asteriadæ
proper. Around the mouth of the Encrinites and Pentacrinites
stand long, ramified, and likewise articulated
tentacula, which remind us of the arms of the Acalephæ
and Cephalopoda.
Fundamentally too, in the Star-fishes, the disk only is
the body proper, and the rays are the tentacula, which,
being monstrously developed, surround the mouth; what
have been called pedicles upon these, represent the suckers
of the Sepiæ.
Eighth Class.
Branchial, Pedal Animals—Crustacea.
Conchozooid Ancyliozoa.
3507. The Crustacea are Worms with horny rings to
the body, jointed feet and tentacula, which mostly breathe
by branchiæ.
They correspond to the Conchozoa, or Shell-animals,
and divide therefore into three orders, each of these
including six families.
Here belong the Crabs, Wood-lice, and Spiders.
In the lowest forms the head, thorax, and abdomen
are blended together, and the back mostly covered with
a great horny scute—Mussel-like Insecta.
In the next place the thorax and abdomen admit of
being clearly distinguished, both by their form as well
as by their appendages, which are much larger on the
thorax, as in the Cray-fish.
Then follow cylindrical forms with numerous feet,
appended to uniform rings, but having a head free and
distinguished by tentacula, maxillæ, and eyes—Wood-lice.
The Wood-lice bear a resemblance to the air-breathing
Insects, although all parts of the body still pass over or
blend gradually into each other.
In the Arachnida or Spiders aerial respiration takes
place, and their body enters into relations with the
more highly organized Insects, since the head becomes
small, the abdomen large, thick, and short.
As the Crustacea pass parallel to the Conchozoa or
Shell-animals, so do they divide into three cohorts or
groups: the Mussel-, Snail-, and Kracken-like Crustacea,
or Crabs, Wood-lice, and Spiders.
Cohort I. Mussel-like Crustacea—Crabs.
3508. Head and thorax connate, being mostly covered
with a shell or shield, abdomen stunted or caudiform;
maxillæ and branchiæ present.
Order 1. Lochmuschelartige, Pfriemenkrebse.
3509. The feet mostly simple and pointed, without
large claws or forcipes, and furnished with setæ or lamelliform
branchiæ; eyes sessile.
Fam. 1. Pholadoid, Mussel-Insects.
Small, almost microscopical, and slightly annulated
animals with an uni-or bi-valvular dorsal shield or testa,
stunted maxillæ, and few legs, having attached to them
setiform branchiæ.
These animals are found in all stagnant waters, in
which they are incessantly paddling or rowing themselves
about. They remind us of those Infusoria which, like
Brachionus, are covered by a scute or shield. They
are what have been called Monoculi (Entomostraca or
Lophyropoda), e. g. Daphniæ.
Number of feet small, and beyond their appended setæ
there are no branchiæ; two eyes frequently blended together;
antennæ mostly furcate like the feet; sexes separate;
a perfect circulation within a true Mussel-like
ventral cavity. They are microscopic Mussels with eyes
and feet.
Fam. 2. Tellinoid, Branchiopoda.
Similar animalcules to the above, but with a body
strongly annulated, naked or covered with a double shell,
and provided with numerous feet, unto which are attached
leaflets that serve as branchiæ.
Here belong the Branchiopoda properly so called.
Fam. 3. Cardiaceoid, Shield-Crabs.
Large animals, with strongly annulated body and
numerous feet; head and back covered by a shield,
whereupon are the eyes—Phyllopoda.
Here range the Trilobites, Apus, and the Molucca or
King-Crabs.
Order 2. Spaltmuschelartige, Scheerenkrebse.
3510. Five pairs of thoracic feet, of which the first
pair is mostly large and forcipiform; the eyes upon
moveable pedicles or foot-stalks.
These animals usually attain a striking size, and are
generally the largest among the horny Ancyliozoa.
The sexual parts open mostly upon the posterior legs.
The abdomen or tail usually supports five pairs of
stunted feet, to which the ova hang.
The number of maxillæ agrees tolerably well with that
of the thoracic feet.
Fam. 4. Mytiloid, Locust-Crabs.
All the feet of pretty equal length, and the forceps
stunted in size; the branchial plates free upon the
abdominal feet—Squilla.
Fam. 5. Arcaceous, Macroura.
Abdominal feet arrested, branchial comb upon the
femora of the thoracic feet and beneath the dorsal shield,
tail extended, as in the common Cray-fish.
Fam. 6. Oyster-like Crustacea, Brachyura.
Characters similar, but tail bent under the body—Taschenkrebse.
COHORT II. SNAIL-LIKE CRUSTACEA—ASSELN.
3511. Body mostly cylindrical, annulate, without a
true carapax or scute; head free; feet short and simple;
branchiæ vesici-or folii-form; mostly placed under the
tail; eyes non-pediculated, or even wanting.
Order 3. Androgynoid—Saugasseln.
3512. Body soft and slightly annulate, maxillæ, feet
and branchiæ arrested; live by suction as parasites upon
other animals, mostly fishes.
Fam. 1. Tritonia-like, Lernæaceæ.
Body soft, without shield, eyes and branchiæ; feet
and maxillæ stunted; ova carried in two tubes at the
hinder part of the body—Kiemenwürmer or Lerneæ.
Fam. 2. Patella-like, Argulaceæ.
Body distinguished into head, thorax, and abdomen,
with few natatory feet, maxillæ moulded to form suctorial
organs, mostly eyes; lay also strings of ova; head elongated
in many in a scutiform manner. The so-called
Fish-lice or Argulaceæ.
Fam. 3. Luftschneckenartige—Pycnogonidæ.
Body short, with four pairs of long feet; eyes, but no
branchiæ and maxillæ; abdomen puny in size. The
Whale-lice.
Order 4. Kammschneckenartige—Nagasseln.
3513. Body cylindrical, horny, and distinctly annulate,
maxillæ, eyes, and mostly seven simple pairs of
feet, branchiæ as cysts or leaves.
Fam. 4. Capulidenartige, Walzenasseln.
Body cylindrical, with five or seven pairs of feet, and
some branchial vesicles; body and abdomen very puny.
The Læmodipoda; Caprella, Cyamus.
Fam. 5. Turbinidenartige, Seitenasseln.
Body horny and distinctly ringed, mostly compressed,
with perfect maxillæ; seven pairs of thoracic feet and
branchial vesicles; abdominal feet rudder-shaped. They
swim usually lying upon the side; many leap—Amphipoda;
Flohkrebse or Gammarinæ.
Fam. 6. Buccinidenartige, Sohlenasseln.
Similar to the preceding family, but the body is depressed,
and the abdominal feet furnished with branchial
plates. The Isopoda; Oniscidæ, to which belongs the
genus Armadillo.
COHORT III. KRACKEN-CRUSTACEA—KOBE.
3514. Body not tripartite; spiracula or air-openings;
more than three pairs of feet, no wings. Here belong
the air-breathing Crustacea; Scolopendræ, Acari, Scorpions,
and Spiders.
These animals are abruptly distinguished from the
preceding by a conversion of the branchiæ into spiral-shaped
tracheæ, which ramify and traverse the whole
body. They all therefore live in the air, and if they do
also dwell in the water, they still come to the surface to
inhale that element. The eyes are only simple points or
ocelli, which are accumulated frequently upon the sides
of the head.
The most inferior of them are distinguished from the
preceding cohort, or the Asseln, by almost nothing
save the essential character of their own cohort, the tracheæ.
They have mostly a number of feet, and only
simple eyes, as the Scolopendræ.
The following have a short body, in which the abdomen
predominates; thorax and head connate; never more
than four pairs of feet—Acari, Scorpions, and Spiders.
They likewise divide into two orders, like the Kracken.
Order 1. Rumpfkrackenartige—Langkobe.
3515. Body horny, tolerably cylindrical and uniform;
feet mostly very numerous.
Fam. 1. Salpoid, Spindelkobe.
Only three pairs of thoracic feet, but still podoidal
appendages to the abdomen—Podura, Lepisma.
Fam. 2. Ascidioid, Schnurkobe.
Body cylindrical, with very many feet; sexual parts on
the thorax—Juli.
Fam. 3. Cirrhopodoid, Bandkobe.
Body band-shaped and depressed, maxillæ perforate,
sexual parts placed posteriorly—Scolopendræ.
Order 2. Kopfkrackenartige—Kurzkobe.
3516. Body thick, mostly globiform; head and thorax
blended together; only four pairs of feet.
Fam. 4. Brachiopodoid, Acarides.
Body rounded; all three parts confluent; usually only
two simple eyes. The Acari are mostly so small, that
their parts can only be distinctly seen through the
microscope. Their mouth is always very much arrested,
and has maxillæ, which are in some cases adapted for
manducation, in others for suction.
Fam. 5. Pteropodoid, Scorpions.
Body tolerably cylindrical, and all three parts connate;
palpi very large and forcipiform.
Fam. 6. Sepioid, Spiders.
Body rounded, head and thorax connate, abdomen
separate, mostly eight simple eyes.
Their most remarkable organs are the four spinnerets
in front of the anus, which probably stand in the signification
of the renal organs, just as the material of the
Spider-threads does in that of the urine.
The tracheæ are but few in number, and expand into
lung-like vesicles.
The sexual parts do not lie posteriorly by the anus,
but at the root or base of the abdomen.
It is moreover remarkable, that their mandibles are
perforated and pour a poisonous juice into the wounds
they inflict. They must be therefore regarded, like the
venom-teeth of Serpents, as elongated salivary ducts.
The Crustacean families can also be named according
to the orders of their circle, namely, the Worms and
Crustacea, e. g. thus:
Order I. Worm-Crabs—Pfriemenkrebse.
Fam. 1. Entozooid—Mussel-Insects.
2. Hirudinous—Branchiopoda.
3. Holothurioid—Shield-Crabs.
Order II. Crustacean Crabs—Scheerenkrebse.
Fam. 4. Typical Crustacea—Sea-Mantes.
5. Asselkrebse " —Macrura.
6. Arachnoid " —Brachyura.
Ninth Class.
Tracheal, Alary Animals—Flies.
Typical Ancyliozoa.
3517. Body tripartite, only three pairs of thoracic
legs, spiracula and wings. They divide, according to
the classes of their circle, into three cohorts, of Worm-,
Crab-like, and perfect Flies.
As the wings are their characteristic organ, so also
must they be divided according to them, and not after
the cibarial instruments. Those wings which are developed
to the least degree are the homogeneous, transparent
wings traversed by few tracheæ or ducts—Tracheoptera,
as Flies, Bees, and Butterflies.
Their chrysalis or pupa condition is perfect.
Then follow wings having very numerous veins united
by transverse veins so as to form a network—Dictyoptera,
as Neuropterous, Orthopterous, and Hemipterous Insects.
The pupa is moveable.
Finally, the anterior and posterior pair of wings become
wholly dissimilar to each other, the former being horny,
and the latter membranous in texture with reticulated
veins, provided likewise with joints like the legs, so that
they can be folded up under the anterior pair or elytra—Ceratoptera,
as the Beetles.
Pupa state perfect.
Strictly speaking, it is a matter of indifference whether
the names of orders and families be adopted from the
first or second parallel series; whether e. g. we speak in
the first cohort of Protozooid or Worm-flies. The
nearest series will, however, bear the greater amount of
resemblance to them. Meanwhile I will in the sequel
vary in the choice of names, in order to exhibit different
samples of this double parallelism.
COHORT I. WORM-FLIES—TRACHEOPTERA.
3518. Wings membranous with few longitudinal
ducts, and almost devoid of transverse ducts, eyes larger
than head. Here belong the Flies, Bees, and Butterflies.
The abdomen is indeed annulate, but soft; its first ring
is frequently set free and unites with the thorax, but supports
neither feet nor wings.
The sexual parts always lie at the anal extremity.
The head is almost nothing but eye, and the Insects of
this order may very well be termed Megalopidæ, or large-eyed,
out of contrast to the succeeding ones. Between
the two large compound eyes there are usually found
three simple ocular puncta or ocelli, which they have
adopted from the preceding class.
The larvæ are either entirely apodal, white and soft
like Entozoa, or they have, in addition to the thoracic
feet, numerous abdominal feet like the higher Worms.
The Flies with their soft and imperfect body, and
the apodal larvæ, repeat the Infusoria and Entozoa; the
Bees therefore the Polyps and Red-blooded Worms; the
Butterflies with their large farinose or dusty wings and
polypodal caterpillars, the Acalephæ and Holothuriæ.
Order 1. Entozooid Flies—Diptera.
3519. All the thoracic rings coalesced, and along with
them the first or basi-abdominal ring, unto which are
attached the halteres or balancers; only the two posterior
wings present; the labium prolonged into a proboscis, which
incloses the setiform maxillæ. Larvæ apodal and white.
The Diptera resemble the Entozoa in a striking manner
through their apodal, soft and white larvæ, and even
through their habitation, which is mostly in fetid and
moist animal matters. Added to this, they respire
usually through two tubes which open upon the anus.
Many during their metamorphosis do not shed their
skin; but it becomes, during the pupa condition, only
horny, representing a small case, whose bottom springs
up in front like an operculum, and gives exit to the
perfect Fly which has been therein developed.
Upon the first abdominal ring of the imago two
nodose pedicles stand out, which are called halteres;
they are probably the old respiratory tubes.
The maxillæ have changed into bristles, which act like
pestels within the groove-shaped labium, puncture and
pump in the fluid.
They divide into three families like the White-blooded
Worms, or according to the orders of their cohort.
Fam. 1. Typical Diptera, Schmeissen.
Antennæ triarticulate, the last joint being mostly
spatulate and furnished with a lateral awn or bristle;
only two suctorial setæ inclosed between the terminal
valves of a fleshy proboscis, which admits of being retracted
within a large cephalic or oral cavity. The
Muscidæ, unto whom the Common House-flies belong,
and also the Hippoboscidæ or Louse-flies. It is my
opinion too, that the Fleas also are to be ranged among
the Diptera.
Alliance 1. Klappenmucken. The two punctuating
setæ placed between two valves without a proboscis—Flea
(Pulex) and Louse-fly (Hippobosca.)
Alliance 2. Acalyptera. Proboscis having thick lips,
and retractile into a large cephalic cavity; alulæ or
halteral opercula arrested—Hypocera to Dolichocera.
Alliance 3. Dung-flies. Characters similar to the
above, but the alulets are of considerable size—Muscidæ.
Fam. 2. Hymenopteroid Diptera, Dasseln.
Antennæ as in preceding family, but the bisetaceous
proboscis is thin and horny, with small labia, or else
large labia with four setæ—Œstridæ, Conopidæ, and
Syrphidæ.
Alliance 4. Parasitic Diptera. Two setæ with or
without a lipless proboscis—Œstridæ, Myopariæ, and
Conopidæ.
Alliance 5. Syrphidæ. Four setæ within a similar kind
of proboscis; the third joint of the antennæ spatulate;
the palpi thickened.
Alliance 6. Leptidæ. Four setæ within a short thick-lipped
proboscis, the third antennal joint mostly coniform—Therexidæ,
Leptidæ, and Dolichopidæ.
Fam. 3. Lepidopteroid Diptera, Gölsen.
Antennæ multiarticulate and stipiform; proboscis with
and without labia, mostly four and six setæ for punctuation—Tipulidæ,
Tanystomidæ, Stratiomydæ, Tabanidæ.
Alliance 7. Gnats. Antennæ filamentary and multi-articular.
Proboscis varied.
Alliance 8. Spiessmucken. Four setæ within a hastate
horny proboscis with or without arrested labia; third
antennal joint not annulated, with the bristle or awn at
the extremity—Tanystomidæ, as Asilidæ, Empidæ,
Bombylidæ.
Alliance 9. Stielmucken. Four or six setæ within a
thick-lipped proboscis; third antennal joint stipiform and
annulated—Stratiomydæ and Tabanidæ.
Order 2. Leech-flies—Hymenoptera.
3520. Four naked veined wings, labium mostly elongated,
the two pairs of maxillæ acting above it like
spears. Larvæ mostly apodal, or with more abdominal
feet or prolegs than in the true Caterpillars.
Most of the larvæ still bear a great resemblance to
the Entozoa, though they do not respire like them
through the anus, but by lateral apertures or spiracula;
the larvæ with abdominal feet repeat the Nereidæ and
Aselli. In other respects they live no longer in putrid
moisture, fungi, roots, and such like bodies, but in living
animals or in cavities specially prepared for their reception
by the parents, and even in a state of freedom upon
leaves.
And here is particularly worthy of our notice the
structure of the cells, which are by many species fabricated
quite substantially of wax or wood-shavings, and
are to be compared with the webs of Spiders, since both
serve as nests for the young. They repeat the Polypidoms.
Others make cases of leaves, and carry into them honey
upon which to deposit their ova.
Others again simply bore holes in wood or in the
earth, in order that they may in a similar manner provide
their young with honey or larvæ. Finally, others simply
stick, by means of their ovipositor, the ova into animals
or leaves.
The dwellings are fabricated by the maxillæ, which
but seldom serve as cibarial instruments, seeing that the
labium undertakes this office as a lick-organ.
Another remarkable feature of this order is the arrested
development of the sexual parts of the females in certain
generations, an occurrence which depends upon the time
of the year or the size of the cells, and whereby they are
constrained to lead a social life as workers.
They divide, according to the Red-blooded Worms or
the orders of their cohort, into three families.
Fam. 1. Dipteroid Hymenoptera, Bees.
Abdomen aculeate; labium elongated in a rostriform
manner. They dig or build cells, and carry thither honey
to the larvæ, which are apodal.
Alliance 1. Fossorial Bees—Andrenidæ.
2. Carpenter-Bees—Anthophoridæ.
3. Cell-Bees—Apidæ.
Fam. 2. Typical Hymenoptera, Wasps.
Aculeate; labium not elongated—rapacious Hymenoptera,
such as Ants, Fossorial, and Cell-wasps, larvæ
apodal.
Alliance 1. Hohlenwespen—Formicidæ.
2. Fossorial Wasps—Sphegidæ.
3. Cell-Wasps—Vespidæ.
Fam. 3. Lepidopteroid Hymenoptera, Terebrantia.
Instead of the sting there is an ovipositor, with which
the females stick their eggs into other insects, mostly
into caterpillars, or else into leaves and wood—Ichneumonidæ,
Tenthredinidæ, and Uroceridæ. The larvæ of
the first are apodal, those of the second provided with
thoracic and abdominal feet like caterpillars, those of the
third with thoracic feet only like the larvæ of Coleoptera.
Alliance 1. Stutzwespen. Ovipositor short; wings
nearly veinless. Deposit their eggs in small insects;
larvæ apodal—Chalcididæ, Oxyuri, Chrysididæ.
Alliance 2. Schlupfwespen. Ovipositor very long,
divided into three hairs; wings veined. Deposit their
eggs in caterpillars; larvæ apodal—Ichneumonidæ,
Evaniidæ.
Alliance 3. Pflanzenwespen. Ovipositor spiral or saw-shaped.
Deposit their ova in plants; larvæ mostly
furnished with feet, and frequently resembling caterpillars
in form—Cynipidæ, Tenthredinidæ, and Uroceridæ.
Order 3. Holothurioid Flies—Lepidoptera.
3521. Four veined wings, covered with small dust-like
scales; maxillæ have coalesced into a proboscis;
larvæ with thoracic and abdominal legs.
The larvæ or Caterpillars remind us of the Nereides,
especially the setaceous Aphrodites, such as the Holothuriæ,
and further still, the Aselli and Scolopendræ. There are
Caterpillars, which are scarcely to be distinguished from
a Wood-louse. They subsist almost throughout upon
leaves, and being exposed to the light are therefore
variously coloured. They have maxillæ, and in the
labium is the orifice of the salivary glands, from which
they spin the threads used in weaving their cocoons, or
social tents. Where they crawl they leave, like the
Spiders, their threads beneath them.
The pupa state is perfect; the perfect insect generally
creeps out of the chrysalis by a slit taking place down
the back.
The abdomen is almost throughout covered with hair,
which is indicative likewise of a strong mucous secretion.
They take in their fluid nourishment by suction, but
the mechanism by which this is effected is not yet
known, as from the jaws themselves forming the proboscis,
there can be no suckers lodged within the latter that
might act. It is probably effected by expansion of the
abdomen taking place during respiration. Their deglutition
would be therefore a respiratory act.
Besides their very non-artistic webs, no artistic instincts
are to be observed in this order. The ova are just layed
without more ado upon plants, and rarely upon other
objects. The Butterflies are generally related to the
plants, and especially to their corollæ, whose colours and
forms they carry in themselves.
They divide according to their proximal orders into
three families.
Fam. 1. Dipteroid Lepidoptera, Moths.
Antennæ filiform, wings mostly thrown like a mantle
round the body; proboscis short.
Small and nocturnal in their habits, proceeding from
tolerably apodal Caterpillars, which reside mostly concealed
in plants, or make themselves cases of hairs and
leaves.
Alliance 1. Typical Moths, Tineidæ.
a. Typical Tineidæ: Alucitidæ.
b. Pyraloid: Tineæ.
c. Tortrix-like: Crambidæ.
Alliance 2. Bombycoid Moths, Pyralidæ.
a. Tinea-like: Aglossæ.
b. Typical: Hydrocampæ.
c. Tortrix-like: Deltoides or Herminiæ.
Alliance 3. Homalopteroid Moths, Tortricidæ.
a. Tinea-like: Fruit-Tortrices.
b. Pyraloid: Heterogeneæ.
c. Typical: Leaf-rolling Tortrices.
Fam. 2. Hymenopteroid Lepidoptera, Silk-Spinners.
Antennæ filiform, wings tile-shaped.
Lepidoptera of considerable size, proceeding out of
Caterpillars, with many feet, and frequently covered with
hair, which live mostly in a state of freedom upon plants,
and in some cases make large cocoons freely exposed, in
others under the earth—Noctuidæ and Bombycidæ.
Alliance 1. Moth-like Spinners, Noctuidæ.
a. Typical: Hadenæ.
b. Pseudobombycoid: Catocalæ.
c. Bombycoid: Erebidæ.
Alliance 2. Typical Spinners, Pseudo-Bombyces.
a. Noctua-like: Psyches, Limacodes.
b. Typical: Notodontæ.
c. Bombycoid: Callimorphæ.
Alliance 3. Homalopteroid Spinners, Bombycidæ.
a. Noctua-like: Bombyces.
b. Pseudo-Bombycoid: Saturniæ.
c. Typical: Hepialidæ.
Fam. 3. Typical Lepidoptera, Homaloptera.
Antennæ varied; wings flat and expanded; proboscis
long—Geometridæ, Vespertina, Diurna.
Large Lepidoptera proceeding from free-living, strongly-coloured,
and mostly naked Caterpillars.
The Geometridæ have level, tolerably triangular wings
and filiform antennæ; they proceed from naked Caterpillars
with few abdominal feet.
The Vespertina have level, long, and narrow wings,
and spindle-shaped antennæ; they emerge from perfect
Caterpillars.
The Diurna have the wings folded upwards in a reverse
position, claviform antennæ, and proceed from perfect
Caterpillars.
Alliance 1. Moth-like Homaloptera, Geometridæ.
a. Typical: Geometræ.
b. Sphingoid: Aposuræ.
c. Papilionaceous: Uraniidæ.
Alliance 2. Bombycoid Homaloptera, Sphingidæ.
a. Geometroid: Zygænidæ and Sesiadæ.
b. Typical: Sphingidæ.
c. Papilionaceous: Castniæ.
Alliance 3. Typical Homaloptera, Papilionidæ.
a. Geometroid: Hesperiidæ, Lycænidæ and Erycinidæ.
b. Sphingoid: Nymphalidæ and Heliconiidæ.
c. Typical: Pieridæ and Papilionidæ.
COHORT II. CRUSTACEOUS FLIES—DICTYOPTERA.
3522. Four wings, with longitudinal and numerous
transverse veins, the anterior pair being mostly leathery in
texture; eyes mostly smaller than head.
Here belong the Neuroptera, Orthoptera, and Hemiptera.
The metamorphosis is tolerably imperfect, and the
larvæ have never more than three pairs of thoracic feet,
and therefore correspond no longer to the Worms, but to
the Crabs, where the number of the feet has been already
more determined. Many also strikingly simulate the
Crustacea.
In all three families also the pupa runs about, eats,
and has rudiments of wings.
The abdominal rings are mostly horny and hairless.
The spiracula lie upon the upper margin.
The eyes are generally much smaller than the head,
and may therefore, in comparison with those of the preceding
and subsequent groups, be said to be of middling
size; the simple eyes have for the greatest part disappeared.
Meanwhile the first family displays large eyes,
the second eyes of a middling size, the third small eyes.
They divide like their antetypes, the Crabs, into three
orders.
Order 4. Crab-Flies—Neuroptera.
3523. All the wings alike and membranous; maxillæ
with mostly large eyes; abdomen soft.
These Insects which, on account of their delicate wings,
are known with us by the name of Flohrfliegen, live
mostly upon flesh, and many of them by capturing living
prey. Many undergo a short pupa stage. Many live in
the water and have, too, branchiæ, whereby they strikingly
remind us of the Crustacea, especially the Branchiopoda.
Like the Crustacea, or in accordance with the two
first cohorts of their class, they divide into two families.
Fam. 1. Tracheopteroid Neuroptera, Nagbolden.
Do not subsist by rapine, but gnaw slowly animal and
vegetable matters; many also as flies eat no longer.
Alliance 1. Dipteroid Neuroptera, Kieferläuse. No
wings; gnaw skin, feathers and hair, mostly Bird-lice—Philopterus,
Liotheum.
Alliance 2. Hymenopteroid Neuroptera, Blumen-and
Mulmlaüse, Thrips, Psocus, Termes.
Alliance 3. Lepidopteroid Neuroptera, Phryganeidæ
(Wassermotten.)
Fam. 2. Dictyopteroid Neuroptera, Raubbolden.
Alliance 4. Typical Neuroptera, Flohrfliegen, Panorpidæ,
Hemerobiidæ, Myrmeleonidæ.
Alliance 5. Orthopteroid Neuroptera, Raphidiidæ,
Mantispidæ.
Alliance 6. Hemipteroid Neuroptera, Ephemeridæ,
Libellulidæ.
Order 5. Oniscal Flies—Orthoptera.
3524. Maxillæ and dissimilar reticular wings; the
anterior, pergamentaceous; abdominal rings, horny; eyes
of moderate size; pupa moveable.
Live for the greatest part on plants.
Fam. 1. Tracheopteroid Orthoptera, Saltatoria.
Alliance 1. Dipteroid Orthoptera, Grylli.
Alliance 2. Hymenopteroid Orthoptera, Locustæ.
Have a hard ovipositor, which reminds us of the sting in
the Hymenoptera.
Alliance 3. Lepidopteroid Orthoptera, Acrydia.
Fam. 2. Dictyopteroid Orthoptera, Cursoria.
Alliance 4. Neuropteroid Orthoptera, Mantidæ.
Alliance 5. Typical Orthoptera, Blattidæ.
Alliance 6. Hemipteroid Orthoptera, Forficulidæ.
Order 6. Arachnoid Flies—Hemiptera or Bugs.
3525. Horny suctorial proboscis with puncturating
setæ; dissimilar reticular wings, the anterior mostly pergamentaceous,
the posterior membranous, and mostly
capable of being folded up, as in the Chafers; eyes small,
pupæ moveable.
The Bugs have much resemblance to the Mites, and
Spiders are as it were winged Mites. One party of them
suck blood, another vegetable sap. They manifest no
artistic instincts.
The Aphides or Leaf-lice require only a single pairing,
to enable them to propagate through several generations.
During the summer they produce only females, and in
autumn the males first appear.
Fam. 1. Tracheopteroid Hemiptera, Leptoptera.
Alliance 1. Dipteroid Hemiptera, Coccidæ, Aphidæ.
Alliance 2. Hymenopteroid Hemiptera, Cicadæ.
Alliance 3. Lepidopteroid Hemiptera, Fulgoridæ.
Fam. 2. Dictyopteroid Hemiptera, Pachyptera.
Alliance 4. Neuropteroid Hemiptera, Water-bugs.
Alliance 5. Orthopteroid Hemiptera, Narrow bugs,
Hydrometræ, Reduvii, Cimicidæ.
Alliance 6. Typical Hemiptera, Broad bugs, Capsidæ,
Lygæidæ, Coreidæ, Scutelleridæ.
COHORT III. PERFECT FLIES—CERATOPTERA, BEETLES.
3526. Maxillæ and dissimilar reticular wings, the
anterior being horny, the posterior membranous and
susceptible of being folded upon themselves; eyes small;
pupæ motionless.
The Chafers or Beetles are the highest Insects, because
they possess the greatest variety of organs,
namely, two kinds of wings; not to mention the perfect
condition of their manducatory apparatus, antennæ,
and the first thoracic ring which is free.
The Beetle represents the Flies in their entire perfection;
everything is rendered horny in the former even to
the lower wings, which are furnished with several joints,
almost like the feet. The upper wings are horny and
meet together so closely by their inner margins upon the
back, that they form a closed suture, like the shells of
the bivalve Mollusc. The antennæ are also more perfect
than in other families, and mostly jointed like the
feet, while at their extremity they are frequently thickened
into laminated moveable clubs, which open when the
animal would fly, as if to listen.
Their habitation, mode of living and subsistence is exceedingly
complex, and therein also they combine again
all the families of this class, and it might be said, all the
preceding classes.
The Beetles are also much more numerous than any
other order, and could in this point of view even hold
good for an entire class, especially, if they represented
a special organic system, but which is not the case.
They live upon vegetable saps, and matters, blossoms,
leaves, and wood, living animals, putrid flesh, dung, and
such like.
They dwell mostly indeed in concealed situations, but
also in those that are freely exposed, while many live in
water, and are so subjected to its influence that their
larvæ actually respire this element through branchiæ.
The larvæ are white and have three pairs of horny
thoracic feet. They live concealed.
The pupæ are invested by a transparent tegument,
which tears in an irregular manner.
The Beetles divide distinctly into three divisions,
which correspond to the three cohorts of this class, or
repeat the three cohorts of their circle.
The Beetles have been pretty generally divided according
to the number of tarsal joints into those having
three, four, five, or an unequal set of these "articuli,"
whereby, however, the greatest disorder has originated.
I have therefore divided them in my 'Naturgeschichte'
according to their mode of living into Phytophaga,
Sarcophaga, and Rypo- or Coprophaga, which gradually
seems to meet with approval. At least a much more
natural arrangement comes to light by using these means.
I have also declared the Rhyncophora to be the lowest
and the Lamellicornes the uppermost in rank, a view,
which at present appears to be generally adopted.
By these means I obtained two firm points, whereby
the division or classification of the intermediate members
is uncommonly facilitated, although many may still stand
in the wrong place. But who can point out a single
system in which Insects range correctly, or, what
is more, wherein it would not be easy to prove that no
single order and family occupies the right place.
In such trumpery systems of classification, one would
think, that some respect might be had for principles, or
at least a feeling of shame for the blockhead's weakness
of intellect. But in vain! Where the sense for philosophy
or for principles is wanting, it is not to be inculcated
or driven in. We still see Natural Histories shoot
forth, with whose shrub-like ramification we must have
compassion. Every thicket of briars is rooted deep, and
admits only of being extirpated by wearisome and patient
culture.
The Rhyncophora indicate that the Phytophagous
beetles are the lowest; the Lamellicornes that the
Coprophagous are the highest. The Sarcophagous chafers
accordingly take their place between the two.
Now, however, the Rhyncophora are tetrameral. In
like manner are the Phytophagous beetles, such as the
Borkenkäfer and Holzschröter. They consequently form
the main stem, unto which all the true Phytophaga must
be annexed. But among them also belong the Pentameral,
namely, the Holzböhrer, Schnell-and Prachtkäfer;
I have therefore disposed them in this order, and I believe
quite correctly.
The Lamellicorn or Dung-beetles are pentameral, and
on that account to be co-arranged with the Necrophaga or
Carrion-feeders and Sexton-beetles, as well as with the
parasitic Lampyridæ, and the predaceous Raub-and
Laufkäfer, despite their exceedingly great difference of
living and even anomalous structure. On the contrary,
most of the heteromeral kinds agree in their mode of
living, and tolerably too in structure, with the Dung-beetles,
at least the Mulmfressenden, such as the Meal-beetles
or Tenebrionidæ, and indeed the Stenelytra also,
while the Cantharides are parasitic at least as larvæ.
It seems also to me, that the fungi must be viewed
as flour or dung, and that therefore those very Fungivora,
whose structure does not point the way otherwise, can
be placed with the Mulm-and Mistkäfern, although they
are only tri-and tetrameral, like the Lycoperdinæ and
Erotyli. Besides this the antennæ of the Fungivora are
usually thickened, and even club shaped, whereby they
thus stand nearer to the Necrophaga, such as the
Speckkäfern and Sexton-beetles, as being those which
likewise live upon putrid substances. It is difficult to
separate the Coccinellidæ from them; their external
resemblance to the Erotyli is also striking.
The pentameral Raub-Lauf-and Sandkäfern form, as
it were, the trunk or main stem of the Zoophagous beetles,
to which are annexed likewise the pentameral Malacopterous
chafers, namely, the Lampyridæ, as parasites.
But the heteromeral Cantharides have also the same
soft wings and mode of life, so that they are not in a
natural system, even if devoid of principles, to be separated
from the Malacopterous beetles.
The nourishment of the Lamellicorn beetles appears
to me to be most perfect in kind. Mould or dung can
be regarded as a fully prepared aliment, or, as it were,
a minced and cooked meat with greens prepared by
Nature, like as Man restores it by art. Thus the lower
Thricozoa, e. g. the Mice, eat the crudest vegetable substances,
such as roots and seeds; those that stand higher,
grass and leaves; then snails, worms, and insects; finally
flesh, and last of all fruits, as the Bears and Apes.
But Man lets the crude matters ferment or reduces them
to rapid decomposition by cooking, whereby a mixed
kind of food results, which bears obviously the greatest
resemblance to dung which, as just observed, is a food
cooked by nature.
According to these considerations I now arrange what
have been called Beetles in the following manner into
divisions drawn from philosophical principles.
The Beetles again commence, like the whole class of
Flies, from below, and the inferior kinds pass therefore
parallel to the Tracheo-and Dictyoptera, while the superior
project above them, as was the case also in the
preceding classes.
In a more remote manner also they repeat the lower
classes, namely, the Protozoa, Conchozoa, Worms and
Crustacea, a fact which is, properly speaking, self-evident,
and which is rendered clear by the following table. We
have thus:
Order I. Tracheopteroid Coleoptera—Phytophaga.
Order II. Dictyopteroid Coleoptera—Zoophaga.
Order III. Ceratopteroid Coleoptera—Rypophaga.
It can also be said; the first correspond to the Worms,
the second to the Crustacea, the third to the Flies.
Lastly, it may still be said; the first correspond to the
Protozoa, the second to the Conchozoa, the third to the
Ancyliozoa.
Order 7. Tracheopteroid Beetles—Phytophaga.
3527. Body cylindrical, head mostly long, antennæ
setiform, maxillary teeth obtuse, tarsi mostly tetrameral.
They gnaw hard seeds, leaves, and wood, and mostly
live concealed. The larvæ almost or entirely apodal.
Fam. 1. Dipteroid Beetles—Rhyncophora.
Head rhynchiform or snout-shaped, tarsi tetrameral.
Fam. 2. Hymenopteroid Beetles, Blattkäfer.
Head tolerably short as well as the antennæ, tarsi
tetrameral.
Fam. 3. Lepidopteroid Beetles, Holzkäfer.
Head pretty short, antennæ very long, tarsi tetra-and
pentameral—Borkenkäfer, Holzbohrer and Schröter.
Order 8. Dictyopteroid Beetles—Zoophaga.
3528. Body long and depressed, antennæ short,
maxillæ large with pointed teeth, feet mostly pentameral
and without ungues.
They subsist upon living or dead animals, dwell usually
in water or upon dry land, and run very swiftly. The
one kind seize upon living beasts, and on that account
constantly swarm about them, as the Raub-and Laufkäfer;
the others only subsist as larvæ by suction upon
other beetles or snails externally, but live, as Flies, upon
leaves and flowers, as the Lampyridæ and Cantharidæ.
Fam. 4. Neuropteroid Beetles—Water-beetles.
Like the following, but all three parts of the body
closely annexed, and rudder-like fringes of hairs upon the
feet—Parnidæ, Hydrophilidæ, Dyticidæ.
Fam. 5. Orthopteroid Beetles—Prædatoria.
Maxillæ very large and pointed, antennæ setiform,
wings hard and tolerably flat like the body, tarsi mostly
pentameral.—Raub-Lauf-and Sandkäfer.
Fam. 6. Hemipteroid Beetles—Parasitica.
Body pretty cylindrical, head rounded with moderate
sized maxillæ, elytra narrow and soft, tarsi penta-and
heteromeral.
They hang as larvæ like Mites to other beetles, and
suck them; but after the metamorphosis they live upon
leaves and flowers—Lampyridæ and Cantharidæ.
Order 9. Ceratopteroid Beetles—Rypophaga.
3529. Body quadripeltate and mostly short, with hard
elytra, thorax large, head small, with short claviform
antennæ and blunt maxillary teeth; feet mostly furnished
with ungues, tarsal joints varied. Their habitation is
very varied, being both free and concealed, and they live
for the greatest part on decaying excrementitious matters,
such as fungi, dry rot, and even animal ordure.
Fam. 7. Phytophagoid—Necrophaga.
Body mostly rounded, with hard and large elytra;
antennæ claviform; tarsi tri-tetra-and pentameral—Lycoperdinæ,
Erotyli, Coccinellidæ, Byrrhidæ, Dermestidæ,
Sylphidæ. They correspond to the Worms and
Tracheopterous insects.
Fam. 8. Zoophagoid—Mulmkäfer.
Body elongated and also nearly spherical, mostly of a
dark spotted colour, with hard elytra, thorax rounded,
head retractile, palpi filiform, tarsal joints heteromeral
or unequal in number—Helopidæ, Diaperidæ, Tenebrionidæ,
and Pimeliariæ.
They live usually in dark situations, eating meal and
dry-rot; have an unpleasant smell. They correspond to
the Crabs, Wood-lice, and Spiders, as likewise to the
Dictyoptera.
Fam. 9. Perfect Beetles—Erdkäfer or Lamellicornes.
Mostly large, convex and short bodied Beetles with
large thorax, broad head, lamellated antennal clubs, spinose
tibiæ and pentameral tarsi. Their habitation is in
dung or under the earth, where they live in the larval
state frequently for years before they change; some
Chafers eat also leaves of trees and flowers. They are
distinguished by their striking size and singular forms,
being mostly furnished with horns on the thorax or
head. I do veritably believe that they may be generally
interpreted as being the highest organized beetles and
members of their class, although they are generally
placed or arranged midway between the lower forms,
while the Sand-and Laufkäfer are esteemed the most
perfect.
Tabular Co-arrangement.
The Dermatozoa now admit of being co-arranged in
the following manner, from which their three Cardinal
relations of Proximity, Repetition, and Serial analogy
or Parallelism, admit of being distinctly recognized.
PARALLELISM OF THE LOWER ANIMALS.
| | | IX Cl. Ptilota. |
| A. PROTOZOA. | VII Cl. Worms. | Co. I. Tracheoptera. |
| I Cl. Infusoria. | O. 1. Weisswürmer. | O. 1. Diptera. |
| F. 1. Monades | F. 1. Saugwürmer | F. 1. Schmeissen |
| 2. Rhizopoda | 2. Tæniæ | 2. Dasseln |
| 3. Rotifera. | 3. Ascarides. | 3. Gölsen. |
| II Cl. Polypi. | O. 2. Rothwürmer. | O. 2. Hymenoptera. |
| F. 1. Hydræ | 4. Earth-worms | F. nbsp; 1. Bees |
| 2. Alcyonia | 5. Nereides | 2. Wasps |
| 3. Actiniæ. | 6. Serpulæ. | 3. Terebrantia. |
| III Cl. Acalephæ. | O. 3. Sternwürmer. | O. 3. Lepidoptera. |
| F. 1. Physaliæ | 7. Holothuriæ | F. 1. Moths |
| 2. Beroeæ | 8. Echinidæ | 2. Spinner |
| 3. Acalephæ. | 9. Asteridæ. | 3. Wanner. |
| B. CONCHOZOA. | VIII Cl. Crustacea. | Co. II. Dictyoptera. |
| IV Cl. Mussels. | Co. 1. Krebse. | O. 4. Neuroptera. |
| O. 1. Röhrenmuscheln. | O. 1. Pfriemenkrebse. | F. 1. Nagbolden. |
| F. 1. Pholades | F. 1. Entomostraca | A. 1. Kieferläuse |
| 2. Tellinidæ | 2. Branchiopoda | 2. Termitidæ |
| 3. Cardiacea. | 3. Phyllopoda. | 3. Phryganeidæ. |
| O. 2. Spaltmuscheln. | O. 2. Scheerenkrebse. | F. 2. Raubbolden. |
| 4. Mytilaceæ | 4. Squillæ | 4. Hemerobiidæ |
| 5. Arcaceæ | 5. Macroura | 5. Raphidiidæ |
| 6. Ostraceæ. | 6. Brachyura | 6. Libellulidæ. |
| V Cl. Snails. | Co. II. Asseln. | O. 5. Orthoptera. |
| O. 1. Androgyni. | O. 3. Saugasseln. | F. 1. Saltatoria. |
| F. 1. Tritoniæ | F. 1. Lernœæ | A. 1. Achetidæ |
| 2. Patellæ | 2. Arguli | 2. Gryllidæ |
| 3. Pulmonea. | 3. Pyncnogonides. | 3 Locustidæ. |
| O. 2. Diœcii. | O. 4. Nagasseln | F. 2. Cursoria. |
| 4. Capulidæ | 4. Læmodipoda | 4. Mantides |
| 5. Turbinidæ | 5. Amphipoda | 5. Blattidæ |
| 6. Buccinidæ. | 6. Isopoda. | 6. Forficulidæ. |
| VI Cl. Kracken. | Co. III. Kobe. | O. 6. Hemiptera. |
| O. 1. Rumpfkracken. | O. 1. Langkobe. | F. 1. Leptoptera. |
| F. 1. Salpæ | F. 1. Poduræ | A. 1. Aphidii |
| 2. Ascidiæ | 2. Juli | 2. Cicadæ |
| 3. Cirripedia. | 3. Scolopendræ. | 3. Fulgoridæ. |
| O. 2. Kopfkracken. | O. 2. Kurzkobe. | F. 2. Pachyptera. |
| 4. Brachiopoda | 4. Acaridæ | 4. Water-bugs |
| 5. Pteropoda | 5. Scorpions | 5. Narrow bugs |
| 6. Cephalopoda. | 6. Spiders. | 6. Broad bugs. |
| C. ANCYLIOZOA. | IX. Cl. Ptilota. | Co. III. Ceratoptera. |
| VII CL. Worms | Co. I. Tracheoptera | O. 1. Phytophaga |
| VIII Cl. Crustacea | II. Dictyoptera | 2. Zoophaga |
| IX Cl. Ptilota or Flies. | III. Ceratoptera. | 3. Moderkäfer. |
| Co. III. Ceratoptera. | O. 3. Rypophaga. |
| A. PROTOZOA. | O. 1. Phytophaga. | F. 7. Necrophaga. | F. 7. Erdkäfer. |
| I Cl. Infusoria. | F. 1. Rhyncophora | A. 1. Fungivora. | A. 1. Phyllophaga. |
| F. 1. Monades | A. 1. Langrüssler | Scaphidiidæ | a. Melolonthidæ |
| 2. Rhizopoda | 2. Kurzrüssler | Erotyli | b. Anoplognathidæ |
| 3. Rotifera. | 3. Breitrüssler. | Encidæ. | c. Rutelidæ. |
| II Cl. Polypi. | F. 2. Blattkäfer. | A. 2. Speckfresser. | 2. Anthobii. |
| F. 1. Hydræ | A. 1. Galerucidæ | 1. Dermestidæ | a. Lepitrichidæ |
| 2. Alcyonia | 2. Chrysomelidæ | 2. Nitidulidæ | b. Dichelidæ |
| 3. Actiniæ. | 3. Crioceridæ. | 3. Silphidæ. | c. Glaphyridæ. |
| III Cl. Acalephæ. | F. 3. Holzkäfer. | A. 1. Dung-feeders. | 3. Melitophaga. |
| F. 1. Physaliæ | 7. Bostrychidæ | 1. Byrrhidæ | a. Cetoniæ |
| 2. Beroeæ | 8. Elateridæ | 2. Sphæridiidæ | b. Trichii |
| 3. Acalephæ. | 9. Cerambycidæ. | 3. Histeridæ. | c. Goliathi. |
| B. CONCHOZOA. | O. 2. Zoophaga. | |
| IV Cl. Mussels. | F. 4. Water-beetles. | F. 8. Mulmkäfer. | A. 2. Coprophaga. |
| O. 1. Röhrenmuscheln. | A. 1. Sumpkäfer. | A. 1. Trachelidæ. | a. Aphodii |
| F. 1. Pholades | a. Heteroceridæ | a. Lagriæ |
| 2. Tellinidæ | b. Parnidæ | b. Pyrochroæ |
| 3. Cardiacea. | c. Hydrophilidæ. | c. Mordellæ. |
| O. 2. Spaltmuscheln. | A. 2. Flusskäfer. | Serropalpidæ. | b. Copridæ. |
| 4. Mytilaceæ | d. Girinidæ | d. Mycteridæ |
| 5. Arcaceæ | e. Haliplidæ | e. Œdemeræ |
| 6. Ostraceæ. | f. Dytiscidæ. | f. Melandryæ. |
| V Cl. Snails. | F. 5. Mordkäfer. | A. 2. Stenelytra. | Arenicolæ. |
| O. 1. Androgyni. | A. 1. Raubkäfer | |
| F. 1. Tritoniæ | a. Pselaphidæ | a. Cistelidæ | a. Æglaliæ. |
| 2. Patellæ | b. Scydmænidæ | b. Helopidæ |
| 3. Pulmonea. | c. Staphylinidæ. | c. Cnodalidæ. |
| O. 2. Diœcii. | A. 2. Laufkäfer. | Taxicornes. | b. Trogidæ. |
| 4. Capulidæ | d. Carabidæ | d. Niliondæ |
| 5. Turbinidæ | e. Brachinidæ | e. Cossyhidæ |
| 6. Buccinidæ. | f. Cicindelidæ. | f. Diaperidæ. |
| VI Cl. Kracken. | F. 6. Schmarotzkäfer. | A. 3. Tenebrionidæ. | Geotrupidæ. |
| O. 1. Rumpfkracken. | A. 1. Leuchtkäfer. | |
| F. 1. Salpæ | a. Lampyridæ | a. Tenebriones | a. Geotrupes. |
| 2. Ascidiæ | b Melyridæ | b. Toxicidæ |
| 3. Cirripedia. | c. Cleridæ. | c. Opatridæ. |
| O. 2. Kopfkracken. | A. 2. Ziehkäfer. | Melanosomata. | b. Lethri. |
| 4. Brachiopoda | d. Notoxidæ | d. Blapsidæ |
| 5. Pteropoda | e. Horiidæ | e. Pimeladiidæ |
| 6. Cephalopoda. | f. Cantharides. | f. Sepidiæ. |
| C. ANCYLIOZOA. | O. 3. Moderkäfer. | F. 9. Erdkäfer. | A. 3. Lohfresser. |
| VII CL. Worms | F. 7. Aaskäfer | A. 1. Phyllophaga | a. Lucanidæ |
| VIII Cl. Crustacea | 8. Mulmkäfer | 2. Coprophaga | b. Passalidæ |
| IX Cl. Ptilota or Flies. | 9. Erdkäfer. | 3. Lohfresser. | c. Oryctidæ. |
Second Province.
— FOURTH CIRCLE—SARCOZOA.
Pisces, Reptilia, Aves and Thricozoa.
3530. These animals necessarily pass through all 4
circles, and each class therefore divides into 4 orders,
or into 5, if the senses be allowed to hold good as
constituting a distinct order in themselves.
Two points of departure for classification here admit
of being thought of, either wholly from below, or first of
all from the commencement of the present circle. In
order to be convinced of this, we need only make a slight
attempt. The animal series is as follows:
| A. Splanchnozoa. |
| Circle I. | Intestinal animals | Protozoa. |
| Class | 1. Gastric animals | Infusoria. |
| 2. Intestinal " | Polypi. |
| 3. Absorbent " | Acalephæ. |
| Circle II. | Vascular animals | Conchozoa. |
| 4. Venous animals | Mussels. |
| 5. Arteriose " | Snails. |
| 6. Cardiac " | Kracken. |
| Circle III. | Respiratory animals | Ancyliozoa. |
| 7. Reticular animals | Worms. |
| 8. Branchial " | Crabs. |
| 9. Tracheal " | Flies. |
| B. Somatozoa. |
| Circle IV. | Sarcozoa. |
| 10. Osseous animals | Fishes. |
| 11. Muscular " | Reptiles. |
| 12. Nervose " | Birds. |
| Circle V. | Aistheseozoa. |
| 13. Sense-animals | Thricozoa. |
3531. Now it is here conceivable, either that the
families of Fishes, Reptiles, Birds, and Thricozoa, pass
parallel to all the preceding classes from the Infusoria
upwards, or that they first commence with the class of
Fishes.
In regard to this question the Birds and Thricozoa
give the most clear and decisive answer. If we adopt
the last proposition, then the Birds must follow each
other thus:
| 1. Fish-like Birds | Palmipedes. |
| 2. Reptilian Birds | Grallæ. |
| 3. Typical Birds | Gallinæ. |
| 4. Thricozooid Birds | Struthionidæ. |
According to this, the Aquatic Birds would occupy
the lowest rank, just as they have actually done hitherto
in all other systems. The three other families too do not
admit of any other position. But what is to be done
meanwhile with the large majority of other Birds? They
would extend or pass beyond the Thricozoa, and have
thus no anatomical system whatever for their basis.
Besides, Humming-birds, Sparrows and Linnets would
in this way rank higher than the Gallinaceous and
Struthious Birds, which some indeed believe to be the
case, but without any proof, so that it is scarcely necessary
to abide by such an opinion.
The Thricozoa must stand thus:
| 1. Icthyoid | Cetacea. |
| 2. Herpetoid | Pachydermata. |
| 3. Ornithoid | Ruminantia. |
| 4. Typical | All the remaining Thricozoa. |
In the systems now in vogue the Thricozoa certainly
follow in this manner, and no scruple whatever is made
of placing the Mice, Ant-eaters, Shrews, and Bats above
Elephants, Horses, and Deer. Yet apart from all this,
the former must pass beyond the Thricozoa, and would
thus likewise have no other organ as a foundation than
perhaps the organs of sense, of which there are but five,
while the families of Thricozoa not yet provided for are
much more numerous, namely, besides those named, the
Seals, Dogs, Bears, and Apes.
The above methods of arrangement must be consequently
cast aside without limitation or reserve, although
they may have held good for hundreds of years. But
what does or will not hold good in an age that is devoid
of principles?
It is thus evident that the smaller Birds stand below
the Aquatics, just as the smaller Thricozoa do beneath
the Whales, quite apart from organic structure, which in
every respect is imperfect. They must correspond, consequently,
to the Asarcous animals, wherewith also their
great number agrees.
If this holds good of the two highest classes, so also
must it hold good of the two lower, namely, Fishes and
Reptiles, and we shall thus have:
| I. | Protozooid Fishes, Reptiles, Birds, and Thricozoa. |
| II. | Conchozooid Fishes, | &c. |
| III. | Ancyliozooid " | &c. |
| IV. | Icthyoid " | &c. |
| V. | Herpetoid " | &c. |
| VI. | Thricozooid " | &c. |
| VII. | Ornithoid " | &c. |
3532. But this parallelism depends essentially upon
the organs, and they are called, in the language of
Physio-philosophy, more correctly:
| I. | Intestinal Fishes, Reptiles, Birds, and Thricozoa. |
| II. | Vascular Fishes, | &c. |
| III. | Pulmonary " | &c. |
| IV. | Osseous " | &c. |
| V. | Muscular " | &c. |
| VI. | Nervose " | &c. |
| VII. | Sensorial " | &c. |
But as these names are unused, and give or convey
only the anatomical idea, but no external image, the
first appellations are to be preferred.
Tenth Class.
Osteozoa, Glossozoa—Pisces.
3533. No one of the upper classes is in such an
extensive state of confusion as that of Fishes. Nor does
this result simply from the great deviations in their
structure from the normal type, these being rather useful
than otherwise for the purposes of classification; but
mainly from the utter want of any principle which might
serve systematists as a guide. Thus at one time we
find them having recourse to the nature of the osseous
tissue, at another to the fins, now to the teeth or to the
scales, ay, even to the fin-rays, and all this because the
characteristic or typical organ has never been sought
after, nor the presence of such an organ as indispensable
been so much as known.
3534. The characteristic organ of Fishes is the Osseous
system, which is consequently the principle also of their
division.
The physical nature, form, position, and number of
the osseous parts must therefore be principally considered,
and hence, above all, the substance or texture of the
bones; the limbs also, and the maxillæ with their teeth, as
well as the teeth upon the palate, upon the tongue and
the branchial arches.
With regard to the component substance or texture of
the bones we encounter a great difficulty. The Cartilaginous
Fishes appear to belong to each other, and are
also usually arranged together. Yet amongst them we
find those species, such as the Lampreys, which obviously
occupy the lowest grade of all Fishes, while the
Sharks and Rays remind us of the Reptilia and Thricozoa,
as well by their external structure as the development of
their sexual parts, since they possess perfect testes, and
ovaria separate or distinct from the oviducts, while they
no longer deposit roe, but large ova inclosed in leathery
shells, like the higher Reptilia. Now, if we separate
these Fishes from the Lampreys, with whom in the scaleless
tegument, the branchial foramina, and even the
external form they have many points of resemblance,
nothing else remains to be done than assign them the
uppermost place, and so parallelize them with the
Thricozoa. But one is next constrained to unite into
one family the Pikes and Herrings, which perhaps admits
of being done.
There belong namely to the upper Fishes without
doubt the Abdominales, which are divisible into five
families: the Carps, Pikes, Shads, Salmons, and Herrings.
Now, if the Sheat-fishes be placed inferiorly on account of
their scaleless body and amorphous maxillæ, four families
will still be left, which should correspond to the Fishes,
Reptiles, Birds, and Thricozoa, so that no place is left
remaining for the Sharks. Now, however, the Salmons
correspond decidedly to the Reptilia; and the Flying
Fishes, which are ranged below the Pikes, probably to
the Birds. If we unite them with the Herrings, then
the Sharks may occupy the place of the Thricozoa.
This being preassumed, we can now attempt the classification.
The substance or texture of the bones is of
such importance, that notwithstanding the separation of
the Sharks and their congeners, the other Cartilaginous
Fishes must be left along with them, and range upon the
lowest stage, so that they thus correspond to the Intestinal
animals.
The next great distinction in the osseous system is the
regular and irregular form which it imparts to the body,
so that the Regular-shaped can be separated in a tolerably
"tranchant" manner from the Irregular Fishes.
The regular form of a Fish is obviously the ellipse, as
we find to be the case in our fresh-water Fishes, namely,
the Perches, Salmons and Carps. They are collectively
covered with large scales, which is therefore also a sign
of their regularity.
The Irregular Fishes are cylindrical, fusi-clavi-spheri-and
tubuliform, usually destitute of scales or covered
with plates, scutes, and spines. Thus, since the Cartilaginous
Fishes are collectively irregular in form, the Irregular
Osseous Fishes must be allowed to follow them.
The greatest variety of the osseous system is shown in
the limbs, especially the posterior pair, which in the
other classes also are generally imperfect and make their
appearance the last. In Fishes they are not divided into
digits, but only into rays, which probably correspond only
to the digital ungues or to feathers. An animal which
has fin-rays is assuredly a Fish, for fin-rays do not occur
in any Reptile.
The posterior fins change even their situation. Those
Fishes in whom they are placed near to the anus, are
obviously the more perfect, as the Abdominales.
In others they advance to the rear of the thoracic fins,
and are even attached to what has been called the "girdle"
or humerus—Thoracici.
In others they even get in front of the thoracic fins on
the throat—Jugulares.
Lastly, they are actually wanting—Apodales.
In the Lampreys there are neither thoracic nor ventral
fins.
The skeleton of Fishes is not simply divided to a
greater extent than in other animals, but has actually a
greater number of bones, such as the rays in the perpendicular
fins, which are wanting in all animals, even in
the Reptilia. An animal with dorsal rays must surely
be placed among the Fishes, and consequently the
Lepidosiren also.
The misshapen Fishes will therefore occupy the lower
stages, the regular the upper, not directly by reason of
their form alone; but because the other organs are also
more imperfectly developed, the bones being cartilaginous,
the tegument asquamous, mucous, or covered with spines,
scutes, and plates; the fins wanting or abortive, or displaced
from their proper situation; the head disproportionate
in size to the body, the mouth wide or narrow,
the eyes placed superiorly or upon the forehead.
A lower character is afforded also by very long dorsal
fins, such namely as extend from the head to the tail.
In the Abdominales, Sturgeons, Sharks, and Rays, the
perpendicular fins are small; in the Thoracici, Jugulares,
and Apodes they are, on the contrary, mostly very long.
Moreover, a lower character is a very long coccygeal
or caudal fin, which denotes that the anus lies far forwards,
and therefore that the tail has a great preponderance
over the trunk. In the Abdominales and the Sharks the
tail is short; in the Rays thin and terminated abruptly,
as in Reptiles and Thricozoa. Thus the higher the
animals ascend the more does the tail diminish in length.
Regard being had to all these relations, the Irregular
Fishes must be viewed as those which correspond to the
lower classes of animals, and the Cartilaginous Fishes
will indeed make the commencement; to these are
annexed the Irregular Osseous Fishes, and nearest to them
indeed those with arrested ventral fins, whether wanting
or placed on the throat; then come the Regular Fishes,
and of these first of all the Thoracici, and next the Abdominales.
In this manner we obtain four divisions.
A. Body irregular.
I. Pisces Cartilaginei.
II. Stummelflosser—Apodales and Jugulares.
B. Body regular.
III. Thoracici—Tunnies, Breams, and Perch.
IV. Abdominales—Carps, Pikes, Salmons and Herrings, Sharks.
Now, these divisions, having been discovered by a
simple analysis or testing of facts, are to be arranged
according to philosophical principles, and further subdivided,
whereby the ground and legality of their existence
will be recognized.
A. IRREGULAR FISHES.
3535. Body deviating from the elliptical form, devoid
of scales, or covered with spines, scutes, and plates;
head and tail disproportionate; fins mostly arrested.
They correspond to the unarticulate Proto-and
Conchozoa; their irregular-shaped mucous or mailed body
agreeing perfectly with these animals.
Order 1. Intestinal, Protozooid Fishes.
3536. Mouth round and without maxillæ, or disproportionately
narrow and wide.
There can be no doubt that the Lampreys are the
lowest Fishes, since they remind us in every respect of
the Worms by their naked, mucous, and lineiform body,
with indistinct head, almost devoid of bones and true
teeth, having a circular mouth, obliterated nostrils, puny
eyes, and finally branchial cysts, which occupy a higher
rank only from their opening into the pharynx. They
pass therefore parallel to the Infusoria, or rather to the
commencements of the second animal series, namely,
the Etozoa Ancyliozoa.
Now, although much dispute prevails concerning the
division of Fishes which should be associated with the
Lampreys, yet it appears to me that no others but the
narrow-mouthed Fistularidæ and Pipe-fishes can follow,
since they resemble them not only in their cartilaginous
bones, but in the structure also of their branchiæ. They
will thus truly occupy the place of the second family.
Whether the narrow-mouthed Globe-fishes are likewise
to be united with them or to be set up as a third family,
may seem to be matter of doubt. I adopt the first
course, and arrange at present the wide-mouthed or
Frog-fishes and Shads in the third family.
Fam. 1. Infusorial Fishes, Lampreys.
Body vermiform, naked, and slimy, without membral
fins. Ex. Branchiostoma or Amphioxys, Myxine,
Petromyzon.
In these Fishes the mouth is quite in front and round,
being without maxillæ and adapted for the purposes of
suction; only one nostril, but mostly several branchial
foramina, which lead to cysts provided with reticular
branchiæ, but without opercula.
The Branchiostomata are the smallest Fishes, not much
above 1" in length, almost devoid of head, yet with
traces of eyes and a nostril. The Myxinæ crawl even
into the rectum of other Fishes, and live therein like
Entozoa. The river or lesser Lampreys stick in the
mud; the Lampreys cling fast by suction to stones, and
do not draw the water in through the mouth, but through
the branchial foramina themselves, like the lower
animals.
Fam. 2. Polypary Fishes, Narrow-mouthed.
Body cartilaginous, mouth having maxillæ, but
unusually narrow, only one branchial foramen with
immoveable operculum—Fistularidæ, Pipe-fishes and
Globe-fishes.
In this family we still meet with species entirely
naked, but covered also with plates, scutes, nails, and
spines. The corymbiform or tufted branchiæ of the
Syngnathi or the Lophobranchii still remind us strongly
of the cystiform reticular branchiæ of the first family.
Fam. 3. Acalephoid Fishes, Wide-mouthed.
Body naked or covered with plates; mouth in front
and mostly unusually wide.
I here arrange in doubt the Frog-fishes and the Shads,
although the last are Abdominales. But they deviate
from the Carps and Pikes by their asymmetrical, naked
or mailed body, the large transverse mouth, the eyes
staring upwards, and the arrested branchial opercula.
Order 2. Vascular, Conchozooid Fishes—Stummelflosser.
3537. Asymmetrical Osseous Fishes, Apodales, and
Jugulares.
Among the true Osseous Fishes the Eels must undoubtedly
range the lowest on account of their vermiform
and asquamous body, and the want of ventral fins. To
them are allied the Jugulares, namely, the Blennii and
Gadidæ, as well as the Plaice; lastly, the asymmetrical
Thoracici without scales, being quite naked or covered
with scutes, as the Gobii and Triglæ.
Fam. 4. Mussel-Fishes, Eels.
Body naked and serpentiform, without ventral fins.
The Eels, from their naked, cylindrical or riband-shaped
body, the long dorsal and coccygeal fin, the small
branchial foramina occasionally confluent beneath the
neck, and by their dwelling in the mud, rank among
the imperfect Fishes. The one set have soft, the other
hard, fin-rays, and on that account they have not simply
been separated, but even far removed from each other.
This difference alone is not so great as to justify their
being arranged, when the structure of the body agrees in
other respects, into distinct orders. The influence of the
dorsal spines upon the life and natatory or waving
movement of Fishes is so slight, and its value generally,
in comparison with other parts, to be taken so little into
account, that a natural arrangement can never result
from these appendages, which do not deserve to be called
organs, and viewed as principal characters.
Fam. 5. Snail-Fishes, Haddocks.
Jugulares without distinct scales and spines on the
branchial operculum, and hard fin-rays.
These Fishes are allied to the Eels by the aborted
ventral fins, the tolerably cylindrical, naked, or small-scaled
body and soft dorsal fins. The viviparous Blennius
bears with us its name of Aalmutter not in vain, for
it resembles a shortened, slimy Eel. The Gadidæ are
indeed less slimy and have in part scales, but, by reason
of their lengthened form and their fins, cannot be
removed far from the Eels; the same holds good of the
Plaice.
Fam. 6. Kracken-Fishes, Grundeln.
Asymmetrical Jugulares and Thoracici, with naked and
mailed body, in addition to hard fin-rays.
Here belong the Gobiidæ and Triglidæ, the first
whereof are usually naked, the second mailed, with roughnesses
upon the head and spines on the opercula; in all
the eyes placed high up.
B. REGULAR FISHES.
3538. Body elliptical, mostly covered with scales, Thoracici
and Abdominales; eyes placed sideways.
Order 3. Pulmonary, Ancyliozooid Fishes.
3539. Regular Thoracici.
The position of the abdominal fins immediately behind
the thoracic is obviously a step further in the perfection
of structure, and these Fishes must be therefore placed
above the preceding kinds, in whom, apart from the
asymmetrical form of body, the position of the fins is
mostly upon the neck or advanced very close to it. The
anus is still situated far forwards, and the tail is therefore
mostly larger than the trunk. The dorsal fins are
still very predominant. Among them we still meet with
naked or microlepidal species, which in their abnormal
form also remind us of the irregular Fishes, as the Tunnies
and Haberdines or Stock-fish. They are therefore to
be regarded as the lowest.
As is the case here in Fishes, so also in the Ancyliozoa
the truly regular or bilateral body originates in them for
the first time, or at least constitutes a persistent character.
Fam. 7. Worm-Fishes, Tunnies.
Tolerably naked or microlepidal, cylindriform or very
much compressed, with small head and mouth; the teeth
very feeble, only like a brush; the branchial opercula
without spines. Here belong the proper Tunnies and
Haberdines: all of them marine Fishes.
Fam. 8. Crustacean Fishes, Brassen.
Body perfectly regular, covered with great scales;
mouth small with strong teeth; branchial opercula unarmed.
Here belong the Labridæ or Lipped Fishes, Seabream
and Osphromanus.
Fam. 9. Ptilotoid Fishes, Perch.
Body quite regular with large scales, mouth of moderate
size with scythe-shaped teeth, branchial opercula
armed. Here belong the Scianoidæ and Percoidæ.
Dwell in the sea and rivers.
Order 4. Sarcose Fishes.
3540. Abdominales, dorsal fins small, with soft ramified
rays, mostly placed far behind. Plainly regular
Fishes with large scales. The head is regular, with the
eyes upon its sides; the set of teeth varied; the trunk
large, tail small, as are also the dorsal fins, which proceed
more and more backwards to the sacrum or even the
tail, a fact indicating their gradual disappearance, and
therefore a sign also of greater perfection. Added to
this, they are generally distributed over the whole earth
in rivers and seas. Finally, it is they that yield the
most nutriment to Man, which is also a constant sign
of greater perfection; as is evidenced in the vegetable
kingdom by the Fruit-trees or plants; in the animal
kingdom by the Oysters, Snails, Sepiæ, Holothuriæ,
Geese, Fowls, Cattle, &c.
Fam. 10. Typical Fishes, Carps.
Body of the Carps covered with large scales, mostly
but one dorsal fin placed pretty far back, mouth nearly
edentate, supra-maxillary bones arrested, large teeth on
the posterior branchial arches, or what have been called
pharyngeal bones. They are for the greatest part fresh-water
Fishes, and those which are most used as articles of
food.
Fam. 11. Reptilian Fishes, Salmons.
Mouth with strong teeth in the superior and intermaxillary
bone; behind the radial dorsal fin there is still
a fatty fin and no large scales. Here belongs the
Lizard-like fish (Saurus), so called on account of its resemblance
to the Lizards. Live in sea and fresh-water.
Fam. 12. Ornithic Fishes, Herrings and Pikes.
The Herrings have teeth in the superior and intermaxillary
bone; only one dorsal fin; mostly large scales.
Body of the Pike slightly scaled, furnished mostly
with a small dorsal fin situated very far back, mouth
full of teeth, but none of these in the rudimental
intermaxillary bone. Dwellers in the sea and in fresh
water.
Among the Pikes is placed the Exocœtus or Flying
fish.
Order 5. Sensorial Fishes.
3541. Bones cartilaginous, mouth opening transversely
under the snout.
Fam. 13. Thricozooid Fishes, Sharks.
Abdominales; bones cartilaginous, mouth opening
transversely under the projecting snout; mostly several
pairs of separate branchial apertures.
Here belong the Chimæræ, Sturgeons, Sharks, and
Rays. The last ought to be held as higher in rank,
partly on account of their slender tail, partly because the
huge Rays, which are called Cephalopterus, have the
anterior thoracic rays free and so moveable that they can
seize their prey with them as with hands. All lay, with
the exception of the Sturgeons, large and leathery ova,
and in this approximate pretty closely to the Reptilia.
It has been already remarked, that the large Cartilaginei
would not pass correctly into the others, and obviously
seem to claim the highest post. At some later
period the principle may probably be discovered whereupon
their union with the higher Osseous fishes depends.
Both are at all events Abdominales.
Eleventh Class.
Myozoa, Rhinozoa—Reptilia.
3542. Body entirely naked or covered with scales,
with distinctly separate and red-coloured muscles; two
nostrils permeable throughout.
As the Muscular system is here the characteristic
organ, it must be regarded principally in the division of
the present class; the limbs also exhibit nowhere so great
a variety as in this class, since they are in some cases
wanting, while in others two only, in some four, are present.
In place of fin-rays true toes have, however, made
their appearance, these again indicating the greatest
variety in the number of the joints; but meanwhile
there are in no instance more than five toes.
The osseous system is constructed after the pattern of
that of the Thricozoa, and is never furnished with dorsal
rays as in Fishes.
The dental formula begins also to be regular. The
teeth stand usually in the superior intermaxillary bone,
and are sometimes pointed, at others obtuse; in the Ichthyosauri
and Crocodiles they are even inserted by gomphosis,
as in the Thricozoa. In many Serpents additional
kinds of teeth occur, namely, the curved poison-teeth or
fangs, which have an involuted groove traversing their
concavity. In most Reptiles teeth also occur upon the
palatal, but there are no longer any upon the lingual bone.
In the Chelonia or Tortoises the teeth are entirely wanting,
and they are also scarcely indicated in the Asquamous
Salamanders and Frogs.
The os quadratum is found as in Birds; in Serpents,
however, the mastoid bone has been freed, and hence the
capability possessed by these creatures of expanding the
mouth.
Viewed in accordance with the perfection of the limbs,
the Salamanders and Frogs ought to be regarded as the
highest in rank; but their scaleless tegument, their
development out of spawn in the water, as well as
branchiæ brings them near to the Fishes. In other
respects their position is determined by the dental formula,
which, as I have already shown, belongs to the
limbs and thus to the motor system.
The position of the nostrils is now, throughout the
class, in front of and upon the snout, being no longer
situated almost on the forehead or vertex, as in Fishes.
Their relation to the scales is likewise of importance for
the purposes of division.
The naked Salamanders and Frogs will occupy the
lowest place. By their form, absence of teeth, and mode
of life, the Chelonia are allied to them.
Then follow the Serpents and Lizards with a perfect
set of teeth. The Serpents are distinguished from the
Lizards by their want of feet and the long bifid tongue
inclosed within a sheath.
Among the Lizards apodal species occur, as the Blind-worms;
but they have under the skin some pedal bones,
and are, in addition to this, sufficiently distinguished from
the Serpents by the short and sheathless tongue.
Among the Lizards with perfect feet there are some
with small visual organs, like as in the Serpents; others
with unusually large eyes, e. g. the Chameleon, Gecko,
Ichthyosauri, and Crocodiles. All other Lizards, the
Serpents, Chelonia, Salamanders and Frogs, have small
eyes and consequently follow each other.
The Crocodile has teeth articulated by gomphosis, and
consequently resembles the Thricozoa.
I divide the present class therefore into the following
groups.
A. DERMAL REPTILES—SMALL-EYED.
Order 1. Protozooid Reptilia-Kröten.
3543. Body slimy and scaleless, or maxillæ devoid of
teeth.
Fam. 1. Infusorial Reptiles, Petromyzoid—Caudate
Batrachia, Salamanders.
Body naked and furnished with a tail.
These cylindrical animals bear the greatest resemblance
to the Lampreys, or are Petromyzoid, being
provided with setaceous and scarcely discernible teeth,
frequently only one pair of feet; they are developed also
out of spawn in the water, and many retain the branchiæ
during the whole of life.
Fam. 2. Polypary Reptilia, Kugelfischartige—Stutzkröten,
Frogs.
Body thick and naked and without a tail, but with
four feet.
The Frogs and Toads proper are likewise developed
out of spawn, but soon lose their branchiæ. In their
form and even the structure of their mouth they are
Kugelfischartige, i. e. remind us strikingly of the Globe-fishes,
among the Engmäulern or Plectognathi.
Fam. 3. Acalephan Reptilia, Welsartige—Chelonia.
Body thick and coated with scales, maxillæ quite
edentulous.
The Chelonia appear certainly to occupy a pretty high
rank; they lay large ova covered with a calcareous shell,
but frequently live or dwell in the water, and have, in
their mode of living as well as form, a striking resemblance
to the naked Batrachia. Through their want of
teeth they range among all the succeeding families.
Order 2. Conchozooid Reptilia—Ophidia.
3544. Body cylindriform and scaly, teeth acuminate,
tongue longitudinally bifid and inclosed within a sheath;
no feet and eyelids.
The Serpents stand without doubt below the Lizards;
yet it is difficult to arrange them properly into families.
The poison-teeth appear to indicate a lower character,
because the cranium is thereby removed or recedes from
the usual type of structure, while the superior maxilla
becomes very much arrested. But as there are moveable
and immoveable poison-teeth, and the latter gradually
pass over into the ordinary kinds of teeth, while externally
also, no character has been found to distinguish
the venomous from the non-venomous Serpents, it is
best at present to discontinue this separation.
Then the Serpents can be brought according to the
structure of their scales into 3 families. They are
either of equal and small size, around the whole body; or
there are plates upon the belly and tail; or finally, the
plates under the latter are divided into two tablets.
Fam. 4. Mussel-Reptiles, Eel-like—Schuppenschlangen.
All the scales around the body small and of equal
size; upon the belly only being somewhat larger. Here
belong the venomous Hydridæ or Sea-snakes, and huge
Boas or Pythons. If once the venomous Serpents could
be distinguished as a distinct family, they will indeed
be placed higher, and the non-venomous be separated
from them. Then probably the Boas may be raised,
together with their allies, as the highest family.
Fam. 5. Snail-Reptiles, Haddock-like—Tafelschlangen.
Plates under the belly, those beneath the tail being
halved or divided into two.
Here belong our Colubri, but also the venomous
Adders.
Fam. 6. Kracken-Reptilia, Grundelartige—Schienenschlangen.
Entire plates upon belly and tail, as in the Rattle-snakes.
Order 3. Ancyliozooid Reptilia—Lizards.
3545. Scales, the usual kind of teeth and tongue;
inferior maxillæ anchylosed in front, mastoid bone not
freed, mostly feet and eyelids.
They divide likewise into three families.
Fam. 7. Worm-Reptiles, Tunny-like—Schleichen.
Body round and serpentiform, with small scales, feet
rudimental, or even wanting.
Here belong the Cæciliæ, Blind-worms, and Scinci.
I have united these animals, which elsewhere have stood
dispersed among the Serpents and Lizards as well as
the Salamanders, and placed them as a distinct family
between the two former. By reason of their rudimental
feet they keep simply upon the earth and bore themselves
passages therein. Their motion is serpent-like,
because they lie upon the belly and cannot assist themselves
with the feet.
Fam. 8. Crustacean Reptiles, Bream-like—Schuppenechsen.
Four perfect feet, body covered all round with small
granular scales, tongue short and hardly slit.
Here belong the Flying Lizards, the Iguanæ and
Basilisks. They usually climb about on trees and seek
for beetles and berries.
Fam. 9. Ptilotoid Reptilia, Perch-like—Schienenechsen.
Four perfect feet, but plates upon the belly and tail,
tongue thin and bifid. Here belong the common Lizards
and the "Sauvegards" or Monitors. Their body is
usually depressed. They cannot climb, but run about
briskly upon the ground and eat beetles, and even the
higher animals. Many are a fathom in length.
B. SARCOSE REPTILIA—LARGE-EYED.
3546. These animals have tubercles, spines and plates,
with four perfect feet, as in the higher Lizards, but the
toes are of pretty equal length; there are no palatal
teeth, and the tongue is not fissured. They are of varied
size, lead a sluggish and mostly nocturnal mode of life,
and have a slow pace, occurring too only in warm
climates.
Order 4. Sarcose Reptiles.
3547. Feet abnormal, being fin-and wing-shaped,
adapted for climbing or clinging. They correspond quite
closely to the three next animal classes.
Fam. 10. Ichthyoid Reptilia, Herring-like—Ichthyosauri.
All four feet converted into fins.
Here belong clearly the monstrous extinct animals,
which formerly lived in the sea. They had gomphotic
teeth almost like the Crocodile.—Ichthyosaurus, Plesiosaurus.
Fam. 11. Typical Reptiles, Salmon-like—Geckos.
Climbing feet, or retractile claws as in the Cats, and
perpendicular laminæ or plates under the soles, by means
of which they cling to walls. They usually dwell in
houses and catch beetles—Chamæleo, Gecko.
Fam. 12. Ornithoid Reptilia, Pike-like—Dragons.
Body short and bird-like, with very long neck and a
long digit, unto which an alary membrane has probably
been attached.
Here belong plainly the fossilized species of a single
genus known by the name of Pterodactylus. They have
been hitherto discovered in different parts of Europe, and
probably flew about, like the Bats, by night.
Order 5. Sensorial Reptiles.
3548. Teeth wedged into the jaws, toes regular.
Fam. 13. Thricozooid Reptilia, Shark-like—Crocodiles.
Feet and toes of equal length, with swimming webs
between them.
The Crocodiles must be without doubt regarded as the
most perfect Reptilia, on account of their notched teeth
and regular feet. In many respects they remind us, like
the Sharks do among Fishes, of the Thricozoa.
Twelfth Class.
Neurozoa, Otozoa—Aves.
3549. In these animals it is again shown in a striking
manner, that the characteristic organ is the principle of
classification. Without recourse being had to the varied
structure of the head, especially of the beak, the division
of Birds would not be possible, although the feet frequently
yield good characters.
3550. The development of the young makes an
essential distinction upon a large scale, since one great
body of Birds must be fed as nestlings, whilst the other,
when scarcely emerged from the egg, runs about and seeks
its own nourishment. Although in the lower animals the
young do not require the assistance of the parents, and,
on the other hand, those of the Thricozoa are suckled
for a longer time by the mother; yet still those Birds
which can, upon issuing from the egg, at once nourish
themselves, such as the Fowls, Geese, &c., are probably
the most perfect, for they pass parallel to the upper
classes of animals, and for the Ostrich to be a Thricozoon,
there is indeed as little wanting as to the Crocodile.
3551. There is no doubt whatever that the Natatores
in every respect, both in the structure of the feet and
head as also in their mode of living and feeding, repeat
the Fishes. This opinion the empirical zoologists have
already adopted from the Physio-philosophy.
The same may be said of the Grallæ or Wading Birds
in reference to the Reptiles. Their feet, neck, and beak
are serpentiform, and associated with a frog-like body.
Their mode of living and feeding is likewise amphibious.
But these birds pass over so directly into the Fowls, that
the latter could not be arranged elsewhere, apart even
from their displaying by their domestication to Man the
higher grade of understanding, which is manifested for
the first time in the class of Birds.
The Bustards and Ostriches are, finally, the highest
stage of Birds, and form the closest alliance with the
Thricozoa.
3552. This point being once settled, it is self-evident
that the Birds, which do not stand in need of being fed,
occupy the uppermost place, and consequently the nestling
or parent-fed Birds the lowest, i. e. the former
correspond to the Sarcozoa, the latter to the Dermatozoa.
A. SPLANCHNIC BIRDS.—NESTHÖCKER.
3553. Remain, after exclusion from the egg, in the
nest and are fed; neck and feet short, toes four in
number and ununited, beak pointed.
By their short neck, which rarely appears longer than
the head, these Birds approximate the Reptiles and
Fishes, in whom also the head has scarcely separated
itself from the thorax.
The uniformity of the feet and toes is likewise an
inferior character, as it is found in the Dermal Reptiles
and Fishes; while in the Sarcose Reptilia and Fishes the
feet and fins make their appearance in the greatest
variety, both as regards the form and length of the feet
themselves, as the structure of the toes; and such is the
case too in the higher organized Birds.
The same holds good also of the form of the head, and
especially of the maxillæ and teeth, which are very
uniform in the lower Fishes and Reptiles, but occur
under very varied conditions in the upper kinds, exactly
as in the Natatores and Grallæ, in the Gallinæ and
Struthionidæ,
Order 1. Protozooid Birds—Tenuirostres.
3554. Bill awl-shaped.
These Birds present a resemblance to each other, not
simply in their mode of feeding, since they are collectively
insectivorous, but also in the slenderness of their form,
and in the dazzling, varied, sharply defined, and very
striking colours of their plumage, as well as in their
habits, for they employ their feet and tail usually for the
purposes of support, and so climb about the upright
stems and branches of trees.
Among them also occur the smallest sized Birds, a
fact which, compared with a similar one in the Thricozoa,
indicates likewise their lower grade or rank.
Fam. 1. Infusorial Birds, Tree-runners or Creepers.
Bill awl-shaped, three toes in front—Humming-birds,
Tree-creepers.
The small size of the Humming-birds seems to render
them the lowest in rank of the class, and by this means
the system obtains a point of departure, unto which
similar forms may be annexed. Their manner of feeding
is rather a process of lapping, than an actual snapping
with the bill; their food also, which consists of small
Beetles and their larvæ, requires scarcely any operation
of the bill, so that here the cibarial instruments obviously
rank upon the lowest stage, and remind us of the proboscis
in Flies, Butterflies, and Bugs.
Fam. 2. Polypary Birds, Woodpeckers.
Bill straight and chisel-shaped, two of the toes directed
forwards and two backwards.
The Woodpeckers stand obviously a step higher,
because their bill is specially active in seeking out larvæ,
and their body is held securely by the toes and stiff tail.
Fam. 3. Acalephan Birds, Cuckoos.
Bill rounded, slightly arched and obtuse; feet scansorial.
These Birds are less animated or lively than the preceding
ones, keep themselves more concealed, and live
mostly upon caterpillars in warmer countries. Some of
them lay their eggs in the nests of other Birds.
Order 2. Conchozooid Birds—Conirostres.
3555. Bill short, thick, and straight, without a notch,
three toes directed forwards.
These Birds usually perch upon branches, without,
however, being able to walk upon them, since they
usually hop, and must therefore make an auxiliary use
of their wings. They are all of them granivorous, and
in a condition both to crush the seeds with their strong
bill, as well as pound them with their fleshy stomach
or gizzard.
Their instruments of manducation and digestion are
therefore perfect, added to which they are so allied to
the subsequent order of Rapaces, that they could not be
arranged lower; probably they ought even to rank higher.
Fam. 4. Mussel-Birds, Sparrows.
Bill short and coniform, without bristles; eat simply
grains.
Fam. 5. Snail-Birds, Crows.
Bill long and coniform; eat grains, beetles and flesh,
and pass gradually over into the Rapaces.
Fam. 6. Kracken-Vögel, Parrots.
Bill very thick and curved. The food consists of
grains and fruits. Their spiritual energies or capacities
are also more strongly developed.
Order 3. Ancyliozooid Birds—Dentirostres.
3556. Bill with a notch; food consists of worms and
flesh.
Fam. 7. Worm-Birds, Cantores or Songsters.
Bill tolerably long and straight; eat worms and
berries.
Fam. 8. Crustacean Birds, Fly-catchers.
Bill straight, with sharp edges superiorly, point de-curved
or bent downwards. Their aliment consists of
insects, which they mostly snap at during flight—Fly-catchers,
Shrikes, Swallows.
Fam. 9. Ptilotoid Birds—Rapaces.
Bill unciform; seize upon the Sarcozoa with their
claws.
B. SARCOSE BIRDS—NESTFLÜCHTER.
3557. Run about soon after being hatched, and nourish
themselves. Bill and feet very varied, the former being
mostly obtuse.
These Birds do not fly much nor hop, but walk, run,
or swim. In them are found all the diversities of bill
and feet; the latter are mostly placed far back, so that
the body is usually directed upwards.
Their food also is very varied, consisting of seeds,
grass, fruits, worms, and flesh.
Order 4. Sarcose Birds.
3558. Neck long, i. e. much longer than the head
and bill, frequently longer than the body.
Fam. 10. Fish-Birds, Natatores.
Natatory feet short.
Mostly large Birds, which live upon fishes, worms,
and many of them even upon herbs. Their bills are so
varied, that they could represent several families, a fact
which likewise speaks in favour of their higher position.
It may be regarded as an instructive hint, that animals
occupy a higher rank, if in them a richness or variety
of forms is to be perceived. This is the case throughout
the Thricozoa. The Natatores, through the structure
and posterad insertion of their feet, through their closely-set
plumage, which frequently presents short and scale-like
feathers, by their swimming and diving, and lastly,
through their fish-catching, approximate as closely to the
structure and mode of living of Fishes as it is possible
for a Bird to do, while still retaining the characters of
its class. Many row even with the wings, and consequently
use them as veritable fins.
Fam. 11. Reptilian Birds, Grallæ or Waders.
Legs, neck, and bill very long.
These Birds are a lively image of, or a composition
from, the Reptilia, having a Frog's body with its long
feet, and a Serpent's neck with a Tortoise's head.
They wade about in marshes to catch worms and
fish; have also bills of very varied structure, yet generally
very long and slender, being in some cases naked,
in others covered with a skin.
Fam. 12. Typical Birds, Gallinæ.
Bill and legs shorter, the former being arched, the
latter armed with strong toes for scraping.
Feed usually upon grains and worms, and live in dry
situations. The Marsh-hens pass at once into the true
Fowls, and these are attached, like as is no other family,
through their capacity for domestication, to Man.
Order 5. Sensorial Birds.
3559. Run only, cannot fly.
Fam. 13. Thricozooid Birds, Bustards.
Fowls with long legs and mostly diminished toes—Bustard,
Cassowary, and Ostrich.
These Birds have mostly such shortened wings, that
they cannot elevate themselves from the earth. In the
Cassowary we find, in place of the primary feathers upon
the wings, five barbless quills like so many claws. The
Ostrich has a pelvis closed in front like the Mammalia.
It has not incorrectly been compared with the Camel,
since the Ruminant animals in general have many resemblances
in common with Birds, especially in the development
of the horny substance, which obviously agrees
with the feathers.
Thirteenth Class.
Aistheseozoa, Opthalmozoa—Thricozoa.
3560. All the senses perfect, five digits; the face
covered with skin and flesh, the body with hairs;
mammæ present.
3561. The Thricozoa combine in themselves all the
animal classes, and indeed, so far as the development by
grades makes no distinction, in equal proportion.
The differences are in no class therefore so numerous
and so strongly pronounced as in this; and yet one is in
more doubt almost concerning the rank of the families
than in the former classes. It turns out, therefore, that
they are not treated according to sound or solid principles,
but that these, after the manner in which Natural History
is still prosecuted, are despised.
Upon viewing superficially the groups of Thricozoa, it
is certainly difficult to determine, which are the lowest
families, although the uppermost ones are very well
known; so that here matters are the reverse of what
they were in Plants. In the case of Birds the empirical
inquirer into nature knows neither which are the upper,
nor which the lower groups; while in the Reptiles and
Fishes he fares still worse.
It is pretty generally conceded that the Whales are
the lowest in point of rank, because their posterior feet
are wanting, and in this they certainly do depart the
most from the usual or normal form of the Thricozoa;
yet still it is just these animals which form the proper
starting-point for the present class. Even the empirical
naturalists here begin to employ physio-philosophical language,
and designate these animals by the title of Fish-like
Thricozoa. But, if they repeat the Fishes, then there
must be Thricozoa, which stand or rank below them, and
pass parallel to the Dermatozoa. Which these latter
are, is not indeed to be declared until the other groups
have been separated and brought into their proper station
or place.
If once there are Ichthyoidal there must also be
Herpetoidal or Reptile-like Thricozoa. Now, if the
Whales correspond to the former, then the serial arrangement
of the latter kinds is readily enough decided. To
the Whales no other animals admit of being annexed but
the Swine-like genera, such as the Hippopotamus, Pig,
and Elephant, which, through their muscular mass, almost
naked skin, and residence in marshy situations, agree very
closely with the Reptilia.
In the next place come the Ruminantia, to take their
site opposite the class of Birds, with whom they correspond
in their susceptibility to domestication, large ears,
fine sense of hearing, and timid disposition. The horns
must be regarded as the obscure metatype (nachregung)
of the feathers. These three families may be aptly termed
Ungulata or Hoofed-animals.
Were matters to fare simply thus, then the center-building
of this class would stand firm, and it would
consequently not be difficult to say which animals stood
below and which above it. For to the Apes are allied
the Bears, to these the Dogs, Cats, and Martens, and to
the latter again the Seals; all would rank above the
Ungulata or Hoofed animals, and represent the proper or
typical class of Thricozoa.
But the Thricozoa are Æsthetic or Sense-animals, and
consequently the upper families must pass parallel to the
organs of sense, if the lower correspond to the anatomical
systems.
Now, if the eye has attained its maximum development
in Man, the same must be said of the ear in the Apes, of
the nose in the Bears, of the tongue in the Carnivora, and
of the skin in the Seals.
Having so disposed of these, the only remaining
Thricozoa are the Bats, Shrews, Moles, Marsupials, Sloths,
and Rodentia.
In common parlance we compare the Bats with the
Birds on account of their wings, unto which may be
further added the large size of their ears. Their close
relation, however, with the Shrews and Rodents assigns
them a lower rank. They must be placed parallel with
the Ptilota or Flies. The dental formula of the Bats resembles
too so strikingly that of the Shrews that a rusty-grown
prejudice can alone place the former in the neighbourhood
of the Apes. They have obtained this post,
without doubt, simply because they have but two mammæ,
and these placed upon the chest.
The Moles cannot be removed from the Shrews.
The choice of position now remains between the Marsupials
and Rodents. And to determine this point cannot
prove difficult, for every one will place the former
above the latter on account of their size, more perfect
dental formula, and the hands, upon whose model the
hind-feet have been in many species formed. To the
Marsupials are annexed the Sloths, because several of
them still possess marsupial or pouch-bones.
Viewed in this and in every other respect, the Rodentia
stand or rank the lowest; and since the members of this
family are much more numerous than those of any other,
we may conclude that they fill up several families of the
asarcose animals.
3562. The series would accordingly be as follows; first
of all Rodents, then Sloths and Marsupials; with Moles,
Shrews, and Bats, all as the repetition of the Dermatozoa.
They are all Myoidal or Mouse-like in character. Next
come the Whales, Pigs, and Ruminants as the repetitions
of the Fishes, Reptiles, and Birds.
The Carnivora with the Bears, Apes, and Man, as
being the proper representatives of the senses, conclude
or wind up the list. They alone have a regular dental
formula. We have thus—
I. Splanchno-Thricozoa; Mice, Edentata, Marsupialia, Shrews and Bats.
II. Sarco-Thricozoa; Whales, Pachyderms, Ruminants.
III. Æsthesio-Thricozoa; Carnivora, Seals, Bears, Apes and Man.
3563. It is here shown, just as distinctly as in the
series of the classes, that no simple scale exists in the
history of development, and consequently in the arrangement
of animals. The Mouse-like species stand off from
the rest, and then follow the entirely different Ungulata
with the Pigs and Ruminants, which once again diverge
in like manner, and make room for the development of
the Seals, which then proceed through the Dogs, &c., in
a less interrupted series up to Man.
He who marvels at this, let him take and set the table
of the class-series before his eyes, and he must give utterance
with us to the following words; namely, that the
lower animals diverge or turn aside, and the entirely different
Fishes, Reptiles, and Birds follow, which, once
again diverging, make room for the development of the
Thricozoa, or, in other words, the "Compendium Animalium."
A perfect parallelism is thus found to exist
between the classes of animals generally and the families
of Thricozoa; but no linear or continuously progressive
connexion is discoverable between one set and the other,
but an appearance by fits or starts of new forms, just as
the systems and organs also are not gradually evolved
metamorphoses of one system, but sudden productions
"en avant" with new tissues, forms, and functions. The
animal system is a multifariously-constructed temple,
with its nave, choir, chapels, and towers, while these again
are present with the whole diversity of forms, which
belongs to them in their several characters or bearings.
A. SPLANCHNO-THRICOZOA—PFOTENTHIERE, MAUSARTIGE.
3564. Small animals with irregular set of teeth; four
feet with claws. The regular set of teeth has included
all kinds of teeth, and with them four or six incisors.
A set of teeth is irregular, which has more or less than
the ordinary number of incisors, in which moreover one
or the other kind of tooth is wanting, or if it is separated
by a breach or interspace.
The small Thricozoa divide into three orders.
The one have blunt uniform molars, two rodent or
gnawing-teeth, and no canines—Rodentia.
The others have, one might say, a wholly aberrant and
confused dental formula, there being at one time too few,
at another too many teeth; molars uniform, with perfectly
irregular incisors and canines—Sloths, Marsupials.
Others, lastly, have a tolerably regular set of teeth,
presenting quadriacuminate molars, mostly small canines,
and rodent-like incisors—Shrews and Bats.
Order 1. Gedärm-, Eyer, Schleimthier-Haarthiere—Rodentia.
3565. Rodent teeth, without canines.
The dental formula of the gnawing Rats and Mice is
so varied, and so devoid of influence upon their bodily
structure and mode of life, that all attempts to arrange
these animals in accordance with it have failed.
3566. The lower position of the Rodentia admits of
being easily proved by taking to our aid the meaning of
the dental system. The splanchnic or visceral teeth
obviously rank lower than the membral teeth. Now, it
so happens that the anterior teeth are in the Rodentia
the principal organ, not only of the dental formula, but of
the entire animal. Without gnawing-teeth the Mice
could not maintain their existence, much less their character.
They thus depend wholly and solely upon the
visceral teeth, and are consequently the lowest Thricozoa.
In addition to this comes their small size, which is by
no means a character to be despised, and one to which all
naturalists pay regard, even while they keep it, upon the
other hand, in the background; or else they must place
the Elephant below the Field-mice.
3567. I divide them, as well according to structure as
habit, into three groups. The one has the front and
hind feet of equal length, with blunt claws for digging.
The other has similar feet, with sharp claws for
climbing.
Lastly, the third has the hind feet longer than the fore
for leaping.
Fam. 1. Infusorial Thricozoa—Wühlmäuse.
Eyes and auditory conchæ feebly developed, feet for
scraping, tail lax; live always under the ground, and eat
roots and grains—Spalax, Rat, Common and Field-mouse,
Beaver; mostly only three molars.
Fam. 2. Polypary Thricozoa—Klettermäuse.
Eyes, ears, and tail large, the latter being stiff and
hairy, claws pointed; live mostly upon trees, eating nut-kernels
and fruit; usually four molars—Squirrels.
Fam. 3. Acalephan Thricozoa—Laufmäuse.
Eyes and ears large, tail hairy, hind feet longest,
claws obtuse, lamellar or plicated teeth; live upon the
surface of the earth and eat grass—Hares and Guinea-pigs;
mostly more than four molar teeth.
Order 2. Ader-, Geschlechts, Schalthier-Haarthiere—Kaumäuse.
3568. Teeth and toes deviating completely from those
of every other order; nor in a less degree the method of
propagation.
In some instances all the teeth are wanting, in some
they exceed the ordinary number, and are quite uniform;
sometimes the lateral teeth only are similar, but in this
case there are no canines and incisors; or else there are
canines with more than six incisors, or also with rodent
teeth.
In like manner the toes are irregular; the one kind
being disproportionately large and the other absurdly
small; some are for the most part wholly connate; in
many cases there are hands posteriorly. The claws also
are in some instances obtuse, in others sharp for climbing,
or else asymmetrically large and unciform. Hands
adapted for swimming or flight also occur.
Finally, the sexual parts are quite aberrant, being
mostly very large and singularly formed; the mammæ
frequently lodged in a pouch, or at least furnished with
marsupial bones. The young are born as naked and immoveable
embryos, and suckled for a very long time.
All this reminds us of the Conchozoa or Shell-animals
as doth also their unusual covering of belts, scales, spines
and long hairs.
Fam. 4. Mussel-like Thricozoa, Sloths.
Lateral and canine teeth equal in size and obtuse, incisors
mostly wanting, and occasionally all the teeth; claws
very large and curved—Ornithorynchi, Ant-eaters, Armadillos
and Sloths.
Fam. 5. Snail-like Thricozoa, Herbivorous Marsupials.
Rodent teeth, usually with stunted proximal and
canine teeth, lateral teeth level; toes mostly connate and
very unequal; they live in the Old World upon roots,
grass, and fruit—Wombat, Dasyure, Kangaroo, Opossum.
Fam. 6. Kracken-Haarthiere, Carnivorous Marsupials.
Mostly more than six incisors, triangular molars and
large canines; live in the New World and in Australia,
eating worms, insects, eggs and flesh—Vulpine Phalanger,
Phascogale, Beutelratze.
The abnormal structure of the sexual parts reminds us
of the same relation in the Snails and Kracken.
Order 3. Lungen-, Fell-, Ringelthier-Haarthiere—Raubmäuse.
3569. Molar teeth mostly quadriacuminate, with a
break in the series, canines and pointed incisors, or
rodent teeth with lateral incisors, five toes; live upon
worms and insects.
Fam. 7. Worm-Thricozoa, Moles.
Claws, sharp incisors or rodent teeth, with lateral
incisors or false molars; live exclusively under the earth,
and cast up the mould.
Fam. 8. Crustaceous Thricozoa, Shrews.
Paws, rodent teeth, with small lateral incisors and
canine teeth. Many burrow passages without throwing
up the soil.
Fam. 9. Ptilotal Thricozoa, Bats.
Alary membrane between the feet and anterior digits;
pointed canine and incisor teeth.
Order 4. Sarco-Thricozoa—Ungulata.
3570. Body large and heavy; teeth stunted, molars
uniform, tolerably obtuse; feet fin-or hoof-like; mostly
udders, rarely mammæ.
Fam. 10. Ichthyoid Thricozoa, Whales.
Skin naked; no hind feet; two horizontal caudal fins;
toes of the anterior feet surrounded by a common skin;
no auditory conchs; posteriorly two udders. All live in
the sea.
It is hardly necessary, in speaking of the Whales, to
direct attention to their monstrously-developed osseous
system and large fleshy tongue, as also to their correspondence
with Fishes in the entire form, manner of living,
and imperfect nose. Their head is still confluent
with the neck, the teeth are horny plates of coalesced
hairs, or uniform simple points, as in Fishes. Most of
them have even dorsal fins; the two udders are hardly
separated from the sexual parts.
Fam. 11. Reptilian Thricozoa, Pigs.
Four feet with hoofs; canines and mostly also incisors;
stomach simple, do not ruminate—Nylghau, Pig, Elephant,
Rhinoceros, Horse. These animals love the
marshes, and are through their mode of living, as also
their form, similar to the larger Reptiles; or, in other
words, through their colossal skeleton, with preponderating
muscular mass, they are Myozoa, through the
proboscis or snout, Rhinozoa.
Fam. 12. Ornithic Thricozoa, Ruminants.
Toes bifid, surrounded by a hoof; above there are
rarely incisor and canine teeth; udders behind; stomach
fourfold; they ruminate. The horn-formation indicates
a relationship with feathers; the want of incisor teeth,
large ears, and timidity of disposition that of the family
with the class of Birds.
Both families enter into connexion with the Whales
through the size of their body, the structure of their feet,
form of the head and disposition towards water and mud.
They are quadrupedal Whales, which have come out of
the water, and adopted a manner of living like the Amphibious
Reptiles and Grallatorial Birds.
Order 5. Æsthesio-Thricozoa—Unguiculata.
3571. Here for the first time an equiponderance of
the sensorial organs makes its appearance and along with
it therefore a resemblance between the animals, which is
no more interrupted by such strange or odd forms, as
in the preceding orders.
All of them have divided toes with claws or nails,
and all kinds of teeth with multiacuminate enamelled
molars.
3572. No doubt can exist about the animals belonging
to the present order; they are the Apes, Bears, and
Carnivora generally, as Seals, Cats, Dogs, Martens, and
such like beasts.
I also cherish no doubt concerning the rank of the
two first families, namely, the Apes and Bears, although
they have been separated in a strange manner by the
interposition of the Bats and likewise the Shrews. At
some future time one will not believe that the Bats and
Shrews were once placed next to the Apes.
3573. Doubt, however, may exist concerning the rank
of the Carnivorous or Rapacious animals; so that here
the principles of our philosophy must be brought to bear
in our behalf. I regard then these animals as the highest
representatives of the sensorial organs. By this step
three families at once take their proper positions, viz.:
Man upon the station or rank of the Eye.
The Apes on that of the Ear.
The Bears upon the post of the Nose.
Difficulties are consequently presented in regard only
to the Rapacious animals, but which are removed, so soon
as the three families just named are parallelized with the
others thus:
Bone, Tongue, Whales.
Muscles, Nose, Pigs—Bears.
Nerves, Ear, Cattle—Apes.
Senses, Eye—Man.
Now the Seals range of themselves next the Whales.
There are thus left the Rapacious animals proper,
which as Dermatozoa or Sentient animals must consequently
correspond to the Splanchnozoa. In favour of this,
evidence is afforded not only by the particular use made
of their feet, but their great number also, which can admit
of comparison with no individual family or order.
Now, however, there are three orders of Murine
animals.
1. Rodent Mice—the Rodentia proper.
2. Chewing Mice—the Sloths and Marsupialia.
3. Rapacious Mice—the Moles, Shrews, and Flitter-mice, or Bats.
In like manner do the rapacious Carnivora divide into
three groups.
1. The mostly sneaking and scansorial Martens and
Viverræ.
2. The sneaking and fossorial Skunks, Gluttons, and
Badgers, with soles and blunt claws.
3. The high-legged Digitigrades, as Dogs, Hyænæ,
and Cats.
As I have given up the dental formula in the Rodentia
as a means of division, so now it seems to me that it
must be abandoned also in the Beasts of Prey. The
whole appearance of them and their mode of living, which
is still the main point in view, obviously directs us more
towards consideration of the feet, than of the dental formula.
The Marten or Weasel kind were formerly compared
with the Mice, and called on that account Mustela.
They cannot be regarded otherwise than as the lowest
in rank.
Unto them are obviously annexed the short-legged
Civets, despite their cunoidal set of teeth. Many have
half soles or pads under the feet. With these again the
Fox-like animals, notwithstanding their viverrine dental
formula, enter into alliance. I believe that I have rightly
parted the Badger from the Bears, and rightly done it too
in this place.
The highest are without doubt the Dogs, Hyænas, and
Cats, with their long and upright legs, not to speak of
their mental faculties. We accordingly obtain the following
arrangement.
Fam. 13. Dermal Thricozoa—Carnaria.
All kinds of teeth included; six broad incisors, a longer
canine, two to three small false molars, a large carnivorous
tooth and large Querzahn, and frequently also the
Kornzahn. Mostly five separated toes, with nails resting
upwards, and either sharp or obtuse feet, occasionally
soled. They eat flesh, mostly that of warm-blooded
animals, and kill their prey themselves.
Alliance 1. Rodent-like Unguiculate Animals—Schleicher.
Feet short and oblique, mostly sharp claws without
entire soles.
1. Mole-like Animals—Martens.
Sharp claws without soles, and no Kornzahn—Martens
and Otters.
2. Squirrel-like—Civets.
Sharp, curved, and mostly retractile claws, superiorly
the Kornzahn—Civet, Paradoxurus.
3. Leporine or Hare-like—Ichneumons.
Straight claws with half soles—Ichneumon, Ryzæna.
Alliance 2. Kaumausartige—Fossores.
Large and straight claws with soles, no Kornzahn.
1. Sloth-like Animals—Mephitic Animals.
Dental formula, like that of the Marten—Skunk,
Mydaus.
2. Wombat-like—Gluttons.
Dental formula, like that of Marten, but behind the
carnivorous is a small tubercular tooth which is wider,
being broader than long.
3. Opossum-like—Badgers.
The tuberculous tooth larger than the carnivorous
tooth, and nearly quadrangular.
Alliance 3. Raubmausartige—Digitigrada.
Legs high and upright; no soles.
1. Mole-like—Dogs.
Claws blunt, the Kornzahn both in upper and lower
jaw.
2. Shrew-like—Hyænas.
Claws blunt; no Kornzahn but a small Querzahn.
3. Bat-like—Cats.
Claws sharp, curved, and retractile; no lower tubercular
tooth, but a small upper one.
Fam. 14. Lingual Thricozoa, Seals.
Feet fin-shaped, the hinder pair stretched out, dental
formula complete, but the lateral teeth tolerably even, and
six or four incisors; tongue mostly somewhat slit. They
correspond to the Whales.
Fam. 15. Nasal Thricozoa, Bears.
Nose elongated into a snout, walk upon soles; all kinds
of teeth, of which, however, the carnivorous or tearing
tooth is similar to the grinders, six incisors.
The slower gait, originating from their walking on the
soles of the feet, with the less pointed molars, assign to
these animals a less rapacious habit of living. They
therefore kill no large animals, and are satisfied with
worms and even roots, fruit, and honey. They repeat the
Pigs.
Fam. 16. Nasal Thricozoa, Apes.
The ears begin to acquire the human form, as do even
the teeth; never more than four incisors, but a longer
canine; hands both fore and aft.
These animals live upon fruits and beetles, and are by
their scansorial feet destined to live upon trees. Their
varied, piping, and sonorous cry is a property of the
sense of hearing, and along with it the larynx also usually
obtains a stronger amount of development.
3574. It seems that every family of Thricozoa contains
five genera, and that these accord with the organs of
sense. In many families this relation is at least striking,
e. g.
Among the Pigs the Elephant is obviously characterized
by the nose, the Hippopotamus by the skin, the Pig by the
dental system; and thus by the sense of taste, the Rhinoceros
by large ears, the Horse by the eyes, thus
| 1. Skin | Hippopotamus. |
| 2. Tongue | Pig. |
| 3. Nose | Elephant. |
| 4. Ear | Rhinoceros. |
| 5. Eye | Horse. |
In like manner do the Ruminantia or Cattle become
distinctly marshalled according to the five senses. The
Camel recedes from the others by its simply uplying
hoofs, the Musk-deer by its monstrous canine; the Goat
is distinguished by its fine sense of smell, and the Giraffe
by its large ears, the Ox by the large and beautiful
eye, thus
| 1. Hautrind | Camel. |
| 2. Zungen- | Musk-deer. |
| 3. Nasen- | Goat. |
| 4. Ohren- | Giraffe. |
| 5. Augen- | Ox. |
In the Bears also we are confronted by this principle
of classification.
| The Skin-bear | is the Common Bear. |
| The Tongue- " | is the Raccoon. |
| The Nose- " | is the Nasua. |
| The Ear- " | is the Arctitis. |
| The Eye- " | is the Cercoleptes. |
After this principle had been so strikingly verified in
several families, I proceeded with confidence also to those
which were more difficult, and it resulted that each consists
only of five genera severally distinguished by the
predominance of some one organ of sense. This method
has been carried out throughout my "Allgemeine Naturgeschichte."
Fam. 17. Ophthalmozoa, Man.
Superiorly or in front hands, inferiorly or behind soles.
3575. Here all the senses enter for the first time into
a state of perfect equiponderance or proportion. Skin
naked, and therefore a perfect organ of feeling; feet and
hands differently constructed for progression and manipulation;
tongue and lips fleshy, while the latter have
hitherto been only tegumental; all the kinds of teeth
different, but still very similar, being of equal height
and nearly equal size; nose elevated by its whole length
from the face, and fleshy; ears oval, laid close against the
head and having regular windings or convolutions; eyes
directed forwards, with perfect eyelids, and moveable in
all directions.
3576. Man by the upright walk obtains his character,
namely, that of bodily freedom, for his hind feet take
the place of all the four feet of other animals, by which
means the hands become free and can achieve all other
offices, the feet alone serving to support the body.
He is the only animal that surveys with the axes of
the eyes borne parallel the most extensive horizon. All
animals whose eyes look higher up or above the ground,
as the Horse, Elephant, Ostrich, and such like creatures,
have eyes directed sideways.
3577. With the freedom of the body has been granted
also the freedom of the mind. Man sees everything,
the whole universe, while the animals can only view
individual parts thereof, two of these even invariably
appearing different, so that the images seen by them are
never reduced to unity.
3578. There is only one human family, only one
human genus, and only one species; and this just because
Man is the whole Animal Kingdom.
3579. But yet there are five kinds or varieties of Men,
according with the development of the sensorial organs.
| 1. The Skin-Man | is the Black, African. |
| 2. The Tongue- " | is the Brown, Australian—Malayan. |
| 3. The Nose- " | is the Red, American. |
| 4. The Ear- " | is the Yellow, Asiatic—Mongolian. |
| 5. The Eye- " | is the White, European. |
Co-arrangement.
3580. The classes of Sarcozoa pass or rank parallel to
each other in the following manner:
| Class X. | Class XI. | Class XII. |
| Anatomical | Osteozoa, | Myozoa, | Neurozoa, |
| Systems. | Fishes. | Reptiles. | Birds. |
| I. Intestinal. | Ord. I. | Ord. I. | Ord. I. |
| Stomach—Infusoria | F.1. Lampreys | F.1. Salamanders | F.1. Tree Creepers |
| Intestine—Polypi | 2. Plectognathi | 2. Frogs | 2. Woodpeckers |
| Absorbents—Acalephæ. | 3. Shads. | 3. Tortoises. | 3. Cuckoos. |
| II. Vascular. | Ord. II. | Ord. II. | Ord. II. |
| Veins—Mussels | 4. Eels | 4. Schuppen- | 4. Sparrows |
| Arteries—Snails | 5. Haddocks | 5. Tafel- | 5. Crows |
| Heart—Kracken. | 6. Grundeln. | 6. Schienens- | 6. Parrots. |
| | chlangen. | |
| III. Pulmonary. | Ord. III. | Ord. III. | Ord. III. |
| Skin—Worms. | 7. Tunnies | 7. Schleichen- | 7. Singing-birds |
| Branchiæ—Crabs | 8. Breams | 8. Schuppen- | 8. Fly-catchers |
| Tracheæ—Flies. | 9. Perch. | 9. Schienen- | 9. Hawks. |
| | Echsen. | |
| IV. Flesh. | Ord. IV. | Ord. IV. | Ord. IV. |
| Bones—Fishes | 10. Herrings | 10. Icthyosauri | 10. Geese |
| Muscles—Reptiles | 11. Salmons | 11. Geckos | 11. Herons |
| Nerves—Birds. | 12. Pike. | 12. Pterodactyli. | 12. Fowls. |
| V. Senses. | Ord. V. | Ord. V. | Ord. V. |
| Thricozoa. | 13. Sharks. | 13. Crocodiles. | 13. Bustards. |
| Class XIII. | |
| Anatomical | Sense-animals, | Aisthesio- | Sense-Organs. |
| Systems. | Thricozoa. | Thricozoa. | 1. Tegument. |
| | 13. Mausartige | |
| I. Intestinal. | Ord. I. | 1. Wühlmäuse | I. Ovum. |
| Stomach—Infusoria | F.1. Wühlmäuse | F.1. Martens | Vitellus—Infusoria |
| Intestine—Polypi | 2. Kletter- " | 2. Civets | Albumen—Polypi |
| Absorbents—Acalephæ. | 3. Lauf- " | 3. Ichneumons. | Envelopes—Acalephæ. |
| II. Vascular. | Ord. II. | 2. Kaumäuse | II. Sex. |
| Veins—Mussels | 4. Sloths | 4. Skunks | Spawn—Mussels |
| Arteries—Snails | 5. Beutelhasen | 5. Gluttons | Milt—Snails |
| Heart—Kracken. | 6. Beutelmarder. | 6. Badgers. | Kidneys—Kracken. |
| III. Pulmonary. | Ord. III. | 3. Raubmäuse | III. Limbs. |
| Skin—Worms. | 7. Scheermäuse | 7. Dogs | Papillæ—Worms |
| Branchiæ—Crabs | 8. Shrews | 8. Hyænas | Feet—Crabs |
| Tracheæ—Flies. | 9. Bats. | 9. Cats. | Wings—Flies. |
| IV. Flesh. | Ord. IV. | | IV. Cephalic Senses. |
| Bones—Fishes | 10. Whales | 14. Seals | 2. Tongue—Fishes |
| Muscles—Reptiles | 11. Pigs | 15. Bears | 3. Nose—Reptiles |
| Nerves—Birds. | 12. Ruminants. | 16. Apes. | 4. Ears—Birds |
| V. Senses. | Ord. V. | |
| Thricozoa. | 13. Aisthesio- | 17. Man. | 5. Eyes—Thricozoa. |
| Thricozoa. | |
3581. By the accompanying table the parallelism of
the different families is recognized, as well as their
remaining relationships, both among themselves as also
with the Asarcous animals, if their table at p. 614 be
compared with it, and which could not for want of room
be inserted here.
3582. It is, moreover, proved from this table that the
classes stand one above the other, but yet that each
recommences from below, so that the lower animals of
a higher class are more stunted or rudimental than the
upper ones of a lower class. Thus the Salamanders are
more rudimental, that is, they have organs more imperfect
than the Sharks; the Tree-creepers are more rudimental
than the Crocodiles, the Mice than the Fowls and
Bustards.
Nevertheless these stunted animals stand higher than
those of the lower classes, because they are characterized by
a higher organ.
What holds good of the classes holds good again of
the orders and families also. The lower animal of a
consecutive family is again more rudimental than the
upper one of the antecedent family. Thus the Ornithorynchus
is more rudimental than the Beaver, the
Shrew-mouse more rudimental than the Opossum and
such like creatures.
In the highest families the equiponderance is first
restored, and the lowest Man is still higher than the
uppermost Ape.