We have now taken a general survey of
the earth’s surface, have noted its mountain
heights and its ocean depths, watched the
formation of hills and valleys which is due
to the joint action of atmospheric agents,
running water and ice, and considered briefly
some of the points of interest about climate.
We next pass to that most characteristic
feature of the surface, its clothing of plants.
Except where the surface of the ground
has been artificially sterilised by man, or is
rendered unproductive by ice, by lava, by
a total lack of water, or by the existence of
poisonous salts, it is clothed with vegetation,
and it is the presence of this vegetation
which is its most obvious character.
Here, however, as in other regions of
thought, the geographical standpoint has
only been reached slowly. Man’s habit of
analysis made him study grasses and trees
for long generations before he got back to the
forest and to the grassland as they occur in
nature. Plants as individuals are the province
of the botanist, but those plant groups
which are the expression of the interaction
of climatic factors, soil, and surface relief, are
the concern of the geographer.
When we take a general survey of the
face of the earth from the point of view of
plant geography, we note three main conditions.
In certain regions, alike in the
tropics and in temperate zones, we find that
plants reach their maximum size, combined
with great differentiation of structure, and
the formation of woody stems which offer
great resistance to varying conditions of
climate and weather. Such highly-organised
plants form forests, which still dominate
over a large part of the earth’s surface.
Man’s nearest allies, the anthropoid apes
and the monkeys, are for the most part forest
animals, and the lowest races of men are still
forest dwellers. Where man is a forest
dweller he seems not to reach his full size,
as we see in the case of the pigmies of the
Congo forest, and the negritos of the Philippines,
and he suffers from a chronic insufficiency
of food, which acts as a check both to
his mental and physical development. There
has, therefore, always been war between
evolving man and the giants of the plant
world, a war which has swept the forests
away from many of the more civilised parts
of the globe, and which still continues, though
man’s victory is now so complete that he can
afford to be generous, and give protection to
the remnants of his former foe.
But over parts of the globe the climate,
and especially the amount or distribution
of the rainfall, makes it difficult or impossible
for forests to grow naturally. Here other
types of plants, lower in stature, and evading
rather than facing the problems of winter
cold or summer drought, flourish and form
what we call the grasslands. The grasslands
favour man in several respects. They feed
the animals upon which he depends for food,
for clothing, and for the conveyance of his
person or property, and they offer much less
resistance than the forest to his agricultural
operations. Even the large herbivorous
mammals which in their wild state haunt the
forests, usually leave these at night to feed
upon the grasslands, so that it is the grasslands
which have largely fed man at every
stage of civilisation. The atmospheric conditions
within the woodlands also, the deficient
sunlight, the humidity, and so forth,
seem unfavourable to human development.
Where the conditions are especially unfavourable
to plant life, we find that even the
grassland plants are unable to keep up the
struggle, and diminish in number, losing
their power of forming a complete covering
for the soil, and thus the grassland passes into
desert, whether the hot waterless desert of
low latitudes, or the cold frozen desert of
northern ones.
In the most general sense, then, we may
say that these three formations, woodland,
grassland and desert, divide the surface of
the land among them, and between them
there is constant conflict. The grasslands
are for ever attempting to encroach upon the
woodlands, and in this attempt they have
been assisted, sometimes to too great an extent,
by the operations of man. Similarly the
desert is always striving to encroach upon
the grassland, and in this endeavour it has
been sometimes involuntarily aided by man,
who has also done much voluntarily to reclaim
the desert land for the grasses.
Let us note next the particular conditions
which favour woodland, grassland and desert
respectively. The distribution of plants over
the surface of the earth at large is determined
by a number of factors, by the amount
of heat, by the amount and distribution of
precipitation, by the nature and strength
of the winds, by the characters of the soil,
and so on. But forests occur under the
equator and also far to the north; we have
cold deserts as well as hot ones; there are
extensive grasslands in the Sudan as well
as in the Canadian Far West. This proves
that the varying amounts of heat may be
neglected in considering the cause of the
distribution of the three great plant formations.
Again, the soil is of minor importance,
for different types of forest and of grassland
occur on different types of soils. We are
thus led to the conclusion that it is the precipitation
and the wind which determine the
distribution. To understand the reason for
this we must consider the needs of different
types of plants in the matter of water.
Plants can only take in the mineral constituents
of their food in the form of a solution,
and this solution must be weak, or it
has a poisonous effect. For example, sulphate
of ammonia is a valuable manure, but
if a considerable amount be dissolved in
water and applied to the roots of a growing
plant, death may very likely take place.
It is a necessary consequence of the fact
that plants can only absorb weak solutions
of their food salts, that their roots take in
more water than is actually needed by the
plant. One of the functions of the leaves
is therefore to get rid of surplus water, the
process being called transpiration. Transpiration
takes place faster in a tall plant
like a tree, which grows up into dry layers
of the air, than in a low plant like a grass.
It takes place faster in windy weather than
in calm. Other things being equal it takes
place faster in warm weather than in cold,
and the larger the plant and the more numerous
its leaves the more water is given off,
that is, the more water is returned to the air
from the soil.
The result of all this is that forest trees
require far more water than grassland. It
has been calculated that a beech wood aged
50 to 60 years gives off during the growing
season 354 tons of water per acre, which
illustrates the drying effect of the presence
of the wood. Similarly, the effect of tree-planting
in the marshy regions of France and
Italy, where the soil as a consequence has
dried and the marshes disappeared, shows
how great a demand upon ground water trees
make, as compared with grasses and low
growing herbs.
On the other hand, although trees take an
enormous amount of water from the soil,
they can draw their supplies from a large area.
It is the extremities of the fine branches of
the roots which take in the water, and these
pass deep down into the soil, and spread out
over a vast area. In other words, trees avail
themselves of the water in the deeper layers
of the soil, and can tolerate relatively long
periods of drought, if the surface drying of
the soil does not extend to the deeper layers.
In hot summer weather grasslands brown
and wither long before the trees show any
signs of water-famine.
In consequence, we may say that as long
as the total rainfall of a region is sufficient
to ensure a constant supply of moisture
in the subsoil during the growing season,
trees can thrive, even if little or no rain falls
during this season. On the other hand, drying
winds are very hurtful to trees, especially
if they occur at a period when the tree is
unable, either because of the coldness of the
subsoil, or because of its dryness, to take in
fresh water to replace that which is lost. The
hurtfulness of late frosts is largely due to the
cold suddenly checking root absorption at a
time when the growing parts, acted upon by
the spring winds, are giving out water freely.
Grasses transpire less freely than trees,
but their root system is much shallower
and less well-developed. They depend upon
the water in the upper layers of soil, and
must have frequent, even if gentle, showers
during their growing season, while they are
quite indifferent to drought and even to cutting
winds during their resting period.
A little reflection will show that it results
from these facts that woodland, grassland
and desert do not form a continuous series.
It may quite well be that woodland passes
through scrub into desert without the intervention
of grassland. Right across Europe
there is (or was) a broad belt of forest. Southward
towards the Mediterranean this thins
out into a characteristic form of scrub, called
maquis, whose characters we shall describe
later, and this scrub passes in all directions
into desert land. Here no belt of grassland
intervenes, for the rainless Mediterranean
summer makes the growth of grass virtually
impossible, except where special conditions,
e. g. hills, introduce modifications. Contrasted
with this we have the conditions in
North America where, e. g. in Canada, the
western coast is densely forest-clad, as is also
the eastern region. In journeying eastward
after crossing the Rocky Mountains the forest
dies away into grassland, and the same
thing happens, though more slowly, in a westward
journey. The reason is that in this
case there is a steady diminution of precipitation
on passing to the interior, but what
precipitation remains is, as we have seen,
largely, though not wholly, summer rain, and
is, therefore, sufficient to determine the
growth of grass, though not of trees.
Again, in North Africa the forests of the
Atlas Mountains pass directly, without intervening
grassland, into the Sahara desert,
but to the south of the desert the grassy and
park-like Sudan separates the desert from
the luxuriant tropical forest. In the latter
case, however, it is possible that man’s influence
has counted for something.
On mountains, in whatever latitude, the
conditions are much more uniform, partly
because it is wind, assisted by temperature
variations, which is the dominating factor.
Moisture is usually abundant, but high up
what is called physiological drought occurs;
that is, the temperature is too low for the
plants to be able to absorb the abundant
water.
In ascending any mountain, the following
are the chief changes which occur. The
lower slopes will probably be cultivated.
As we ascend the precipitation increases, and
forests appear. First we have probably a
belt of deciduous trees, passing above into
the more resistant conifers. This belt usually
ascends higher on the south than on the
north side, and higher on mountains which
occur in a group than on isolated peaks. As
the wind is more and more felt, and increases
the dangerous transpiration of winter the
trees become more and more dwarfed to
escape its force. There may be a belt of
prostrate mountain pines above, marking the
tree limit; in any case the trees are gradually
replaced by dwarfed shrubs. Then comes
the zone of Alpine plants, the grasses making
a complete sward, but being accompanied
by many other plants. Gradually, as the
soil becomes scantier, and the surface more
rocky and exposed, the continuous sward
disappears, and the conditions of a cold desert
appear. A few scattered plants occur, ceasing
near the snow-line, the highest being
usually plants of simple structure like mosses
and lichens.
As we have already indicated, in the case
of the mountains of Europe there are often
glacial shelves at considerable elevations,
whose covering of fine débris determines
the growth of peculiarly fine grass. The
economic value of this grassland has in many
cases in the Alps induced man to destroy
the forest in order to increase pasture land.
The result has often been disastrous, for
once the trees are cut down the forest soil is
rapidly destroyed by weathering, especially on
slopes, the courses of streams are altered by
the more rapid run-off, and widespread flooding
and destruction of pastures have sometimes
resulted. In North America, similarly,
man’s attempt to increase pasture land or
arable land at the expense of woodland has
often led to disastrous consequences.
We have already spoken of the special
features of the Mediterranean climate, and
indicated that its peculiarities are reflected
in its vegetation; we must now consider this
vegetation in a little more detail. The fact
that the region is chiefly visited by the
inhabitants of more northern climates in
spring gives rise to a somewhat erroneous
impression in regard to the plants. In spring
the Mediterranean vegetation is at its best.
The mild winters permit the plants which
further north die down or cease to grow, to
go on blooming. The rains so moisten the
soil that the first warm days cause very rapid
growth in those plants which finish their
activities before the hot, dry summer begins.
They must flower and seed in spring, and
die down till the rains of autumn awaken
them again.
In our own country we have a few plants
which hurry through their activities in this
way. The lesser celandine, the wood anemone
and a few others strive to flower and fruit
before the forest trees are thickly clad with
leaves. The snowdrop, even the wild hyacinth,
though it is much later, similarly
limit their active life to a short period in
spring. This phenomenon, only suggested
in our climate, is very marked in the Mediterranean
area.
That region is especially characterised by
its richness in bulbous and tuberous plants.
These, as all who have grown hyacinths or
narcissuses know, demand relatively large
amounts of water during their short growing
period. In spring, therefore, the shores of
the Mediterranean are bright with many
kinds of anemones, with narcissus, asphodel,
bell hyacinth, Allium, tulips, and so on, all
awakened by the spring warmth and the
spring rains. Accompanying them are many
bright-coloured annuals, also in a hurry to
race through their life-history before the
terrible drought of summer. Now also the
grass grows, and the autumn-sown corn becomes
tall. As the weather grows hotter and
drier, the plants with bulbous and tuberous
roots die down to the ground, the annuals die
altogether, leaving their seeds to wait till the
autumn rains before they sprout. The grasses
turn brown, and the peculiar parched appearance
of the Mediterranean summer
spreads over the land.
To a northern visitor at this season it is not
luxuriance but desolation which is the prevailing
note. Except on the hill slopes
there are no masses of broad-leafed foliage
trees—there is not the deep bright green
characteristic of the summer woods further
north. The trees do not reach a great size;
the leaves are usually small, and the fact that
they strive to avoid the sun by arranging
themselves with the edge upwards instead of
the flat surface, makes them appear smaller
than they are. They are often needle-shaped,
sometimes shining and coated with resin,
sometimes silvery owing to a coating of hairs
on the under surface. Many plants have
spines or thorns, and succulent plants like
agave, aloe and prickly pear are common.
The absence of a complete covering of
vegetation causes the surface soil to dry
completely, and so form clouds of dust which
adds to the generally desolate appearance.
Indeed, the brown powdery appearance of
the soil is one of the points which especially
strikes the stranger, accustomed to the darker,
moister soil of the north, always covered
with vegetation, except where man has
interfered.
Here and there, however, are indications
that even this parched brown earth holds
wealth for man. The vines, if dusty and far
less luxuriant than one expects, are loaded
with ripening fruit. The gorgeous scarlet
flowers of the double pomegranate gleam amid
the dark foliage; the gnarled and twisted
olives show on close inspection masses of
small green fruits; the oleander bushes are
covered with pink flowers; there are great
round balls on the orange and lemon trees,
and many other fruit trees are loaded with
produce.
Let us sum up first what man gains from the
plants of the Mediterranean, and then look
at some points in regard to the wild plants.
In the first place, we see that man takes advantage
of the rapid growth of annuals in the
early part of the year. The annuals most
useful to him, here as elsewhere, are, of course,
the cereals, especially wheat, which, if sown
in autumn, is nourished by the winter rains,
and grows rapidly with the warmth of spring
to ripen in May, June or July, according to
the locality.
In the second place, certain trees or shrubs,
by reason of their resistance to drought, and
their elaborate root system, which enables
them to gather water from the deeper layers
of the soil, will produce succulent fruits
without needing artificial supplies of water.
The most important of these, throughout the
whole Mediterranean area, are the vine and
the olive. The olive supplies the oil which
is all the more necessary in that the absence
of grass makes pastoral industries, and therefore
the production of cheese and butter
difficult or impossible except in the high
grounds, while the vine supplies the wine
which with bread and oil form the essential
parts of the diet of Mediterranean man.
The olive tree, which is indigenous, may be
regarded as one of the most characteristic
trees of the area, and it is interesting to note
that the novice not infrequently confuses
it with another tree, almost as characteristic
the evergreen or holm oak. The two are not
nearly related, the olive belonging to the
same family as the lilac and privet, while the
evergreen oak is a true oak. Both trees,
however, show similar adaptations to summer
drought, and their resemblance to one another
is a good example of convergence due
to a similar environment. Both have small
evergreen leaves; small that they may not
lose too much water in summer, evergreen
that they may assimilate even during the
winter. Both have their leaves silvery beneath,
which again prevents loss of water;
both have gnarled trunks, branching low
down, in order that the leaves may avoid the
dry upper layers of the air. Adaptations of
this kind are present to a greater or less degree
in all the trees which are tolerant of
Mediterranean conditions, and many of these
trees yield useful fruits.
In addition to the cultivated plants mentioned,
a great number of others are grown
within the area, as we shall see later, but the
point of interest is that the plants which have
been of importance in the history of the region
have been either annuals which ripened early,
or fruit-bearing trees with special adaptations
to resist drought.
Apart from the annuals and the bulbous
and tuberous plants already described, the
wild plants are chiefly shrubs or stunted trees
with similar drought-resisting characters.
During the long ages he has inhabited the
Mediterranean, man has doubtless contributed
largely to the destruction of the forests
which are now, as we have seen, represented
by the stunted scrub or maquis. But on
climatic grounds we cannot suppose that the
Mediterranean forests had ever the luxuriance
of those further north, or of the tropical
forests of the south.
Where there is sufficient rain chestnut
woods occur, but this is only on the hill slopes.
Above the chestnut, beech may occur, as in
Sicily. The maritime pine and the Corsican
pine form open woods in the damper places,
and the picturesque stone pine, with its
rounded head, is very characteristic. We
have already mentioned the evergreen or
holm oak as common, and the cork oak occurs
abundantly in some places. These trees, with
the cypress, must have formed the primitive
forests, and they still constitute the most
important forest trees of the area. The occurrence
of a native palm (Chamærops) is interesting
as suggesting the warmth of the climate,
and even on the European shores the
date palm is extensively planted, though its
true home is the margin of the African and
Arabian deserts.
Of the characteristic shrubs the most
striking are perhaps the many species of Cistus,
with large almost rose-like flowers, and
leaves which attempt to adapt themselves to
the climate by many different devices. Sometimes
they are stiff and leathery, sometimes
resinous, sometimes hairy. Many plants in
the area have a coating of resin on their leaves.
This, no doubt, preserves them against loss
of water, but also probably protects against
grazing animals. Goats thrive in the
Mediterranean partly because of the catholicity
of their taste in vegetation, and in consequence
the plants have had to protect themselves
against their appetite as well as
against drought. Only those with some disagreeable
quality, hairs, spines, resin, strong
flavour, etc., could hope to protect themselves
in the dry season, when grass is virtually
absent. It is in consequence common to
find aromatic or strongly-flavoured plants
with glandular leaves; lavender, rosemary,
myrtle, etc., are examples.
Other shrubby plants associated with the
Mediterranean are oleander, the noble laurel,
the tree heath, arbutus, many kinds of broom,
and generally evergreen shrubs specially
adapted to resist drought.
Let us turn from this picture to the appearance
presented by Central and Northern
Europe. As we have seen, the forest which
once covered most of the area, except the
steppe region of southern Russia, has largely
disappeared, but enough remains to enable
us to reconstruct the picture of the original
forest.
As contrasted with the (chiefly) evergreen
woodland of the Mediterranean, the forests
of the low grounds are here deciduous. In
summer clothed in magnificent foliage, well
adapted to give off enormous quantities of
water, in winter the trees stand tall and bare,
exposing nothing but their branches to the
winter blasts. While the buds of Mediterranean
plants have no special means of protection,
the typical forest trees of Central
Europe have their buds carefully sheathed
in scales, clothed in hairs, or coated with
resin, to keep out alike the cold and the damp
of the northern winter. While the leaves of
Mediterranean plants are usually small, often
coated with hairs beneath, often resinous,
and so on, the forest trees further north have
large leaves of delicate texture, with no
special protection against drought.
Again, while the luxuriant forest of the
tropics includes many different species of
trees, the deciduous forests of cool temperate
regions contain few species, and are often
pure woods, that is, consist of one dominant
species, forming beech woods or oak woods,
and so on. The dense shade of the beech
makes undergrowth difficult or impossible,
but the other woods have a complicated
undergrowth of many different kinds of
plants, especially pronounced in spring before
the leaves appear on the trees. But this undergrowth
never reaches the luxuriance that
it does in the tropical forest, and creepers and
climbing plants are few.
As we ascend from the low ground to the
higher, or as we travel northwards to high
latitudes, the broad-leafed deciduous forests
are replaced by coniferous ones. European
conifers, with the exception of the larch,
are evergreen, and all are more tolerant of
cold and wind than deciduous trees. Pines,
spruce, fir, larch, and silver fir are the most
important kinds. Both at high altitudes
and in high latitudes these conifers are often
accompanied by birch, which is not a cone-bearing
tree, but is very tolerant of cold and
wind.
To the north there comes sooner or later
a limit beyond which the cold and winds
make further tree growth impossible. Here
we come to a tundra region, where the place of
trees is taken by low-growing shrubs, with
small leaves and other adaptations to ensure
against excessive loss of water. It is, as it
were, the reappearance of the Mediterranean
type, but here the cause is, not the absence of
water, but the fact that the cold makes it
impossible for the roots to absorb it. A condition
of physiological drought results, and
only plants well adapted to prevent undue
loss of water can resist such conditions of
life.
A somewhat similar type of vegetation
occurs over vast areas in the more northern
parts of Europe, forming the moors and
heaths of much of Scotland, of parts of England
and Ireland, of parts of Germany, and so
on. Here the presence of peat produces conditions
very unfavourable to plant life, except
to certain shrubby plants such as heather and
other plants of the heather family, juniper,
bog myrtle, and so on, and some grasses
and sedges, etc., all of which have special
adaptations to life in a peaty soil. Over the
large areas, therefore, covered by these heaths,
trees are absent, or few, and this stunted
shrubby vegetation takes their place.
Large areas of natural grassland, except
for the tracts of pasture land already described
in the mountain regions, are infrequent
in Europe. They occur in Southern
Russia and in the Hungarian plain, and form
part of that great series of steppes and plains
which stretches into Asia, and passes into a
region of deserts.
The conditions favourable to the growth of
grass here, instead of trees, seem to be purely
climatic. Very important is the prevalence
of strong cold winds during winter, which is a
period of drought. The scanty rains come
in early summer, which suits grasses admirably,
while the total precipitation is too slight
for trees. The summers are hot, and the
rains cease early and give place to a period
of drought, very injurious to trees, while it
injures the grasses little, owing to the fact
that they have had time to make their
growth.
The abundant natural growth of grass
makes these steppe regions well suited to the
pastoral industries, which tend, as civilisation
progresses, to give place to agriculture.
To sum up, we have seen that looking at
Europe as a whole three great plant formations
are represented. We have, first, the
cool temperate forest, which once extended
over the greater part of the continent, wherever
the conditions were suitable. This has
now largely given place to arable land. Next,
we find round the Mediterranean sea, and in
those great peninsulas and islands which are
bathed by it, a zone of modified woodland
passing into scrub, remarkable for the rapid
growth of annuals in the early part of the year,
and for the abundance of trees bearing useful
fruits. Finally, linking Europe to temperate
Asia, we have belts of steppe land, characterised
by a luxuriant growth of grass in the
early summer, and fitted by nature for
pastoral industries, which do not thrive near
the Mediterranean. Another way of putting
the same facts would be to say that Europe
proper is a region of temperate forest, linked
to Africa by scrub land passing into desert,
and to Asia by steppe land passing into
desert.
The flora of North America, owing to the
size of the continent, offers more resemblance
to that of Asia than to Europe.
Bearing in mind what has been already
said about the structure of North America—with
its western mountain range and eastern
uplands enclosing between them a region
of moderate relief—and also what has been
said in regard to its climates and to the influence
of climate upon vegetation, it is
relatively easy to deduce the main points
in regard to the flora.
To the far north there is a treeless tundra
region, quite comparable to that which
occurs over vast areas in North Asia, and on
a reduced scale in the northern part of the
continent of Europe. Next we have a wide
band of predominantly coniferous forest,
which, although its species are different, yet
in broad outline is entirely homologous with
the coniferous forest found in northern Asia,
south of the tundra region. In Canada this
forest consists of spruces and larches, the
species being peculiar to the continent.
Mingled with the conifers are smaller numbers
of the hardier deciduous trees, such as
birches, poplars, and willows.
What we have already said as to the
climatic differences between the eastern and
western sides of continents will at once suggest
that this band of forest is not likely to
run directly across the continent from east
to west. In point of fact it stretches from
Labrador in a north-westerly direction to
Alaska, leaving almost the whole of the
western seaboard to be occupied by another
type. This type is the extraordinarily luxuriant
and beautiful western forest, consisting
for the most part of conifers. It is largely
these conifers which have enriched European
parks and gardens within recent years, and
although it is perhaps the great Sequoia
(Wellingtonia) gigantea which has most impressed
popular imagination, it must be
remembered that size and luxuriance are
characteristic of many species. This western
forest stretches down the western seaboard to
the State of California, and, indeed, persists
until increasing aridity makes forest growth
impossible. Its great luxuriance, compared
with the scantier forests of the Mediterranean
region in Europe, is partly to be ascribed
to a greater rainfall, and doubtless partly
to man’s interference, for the original forests
of the Mediterranean must have been largely
destroyed, as the western American forests
are in process of being. One must remember
also that the proximity of mountain ranges
to the seaboard in western North America
gives a heavy rainfall, and suitable places
for forest growth. The fact that the trees are
predominantly coniferous gives them great
resistance to the summer drought. In front
of the mountain ranges the coastal plain is
occupied by an evergreen scrub vegetation
comparable to that of the lowlands of the
Mediterranean basin.
In British Columbia, where the Cascade
Range lies at no great distance from the
Rocky Mountains, the western coniferous
forest practically clothes the whole area from
the coast to the main range, but further south,
where the Cascade Range and its continuation
the Sierra Nevada are widely separated
from the main range, a dry and semi-desert
region occurs, between the two, which bears
a desert type of vegetation, including especially
a plant related to our wormwood, called
sagebrush, with cactuses in the warmer parts.
Another area which is too arid to carry trees,
except where local conditions raise the rainfall,
extends from Texas northwards to about
the latitude of Edmonton or Battleford,
and lies in the “rain shadow” of the Rocky
Mountains. This is the region of the Great
Plains, mostly too arid to carry anything
but herds of cattle, and mostly forming
natural pasture, being thus analogous to the
steppes of Asia.
Eastward the rainfall increases, and we pass
from the area of unreclaimed pasture to the
prairies, now largely laid down to wheat and
other food plants. Southward the Great
Plains pass into the deserts of Mexico, but
northwards they are separated from the
northern coniferous forest by a belt of aspen,
and it is in this region that the Canadians
are steadily pushing the cultivation of wheat
into the plains, wherever the local rainfall
makes this possible.
So far we have left south-eastern Canada
and the whole of the eastern and south-eastern
States out of consideration. Speaking very
broadly, we may say that all this area is
clothed by a forest of mixed coniferous and
broad-leaved trees which is comparable to
the forest which covers the greater part
of temperate Europe. But it is not to be
expected that a forest which extends from
the northern shores of the Gulf of St. Lawrence
to the extremity of the peninsula of
Florida, that is, through about 25 degrees of
latitude, should be uniform throughout. In
point of fact, botanists distinguish three
separate zones. In south-eastern Canada
and the New England states the Weymouth
pine (Pinus strobus) predominates, being
accompanied by limes, ashes, maples, oaks,
elms, chestnuts, and so forth. Further
south, and especially further west, extending
to the Mississippi plains, there is a deciduous
forest extraordinarily rich in species. Practically
all our common genera of forest trees
are represented, sometimes by very fine
species, but in addition there are many
genera with no European representatives.
Very striking is the abundance of magnolias
(whence the name of magnolia forest sometimes
given to this type), and species of the
laurel family, as well as of liquidambar.
The magnolias and liquidambar are especially
interesting, because they once occurred in
Europe, their disappearance there being
probably caused by the glacial period as
explained on p. 78.
We have emphasised above (p. 137) the
luxuriance of the forests of the west coast of
the States, but it should be noticed that
luxuriant as its conifers are, there is a remarkable
poverty in broad-leaved forms, as compared
with these eastern forests, and this
even in the warmer parts of the west coast.
The reason is probably the same as in the
Mediterranean region in Europe. The existence
of a belt of desert to the south of the
present “Mediterranean” region of western
America made it difficult for the trees to
migrate southwards at the onset of cold conditions
in the glacial period, and thus many
forms, which are known to have existed in
California in Tertiary times, have now completely
disappeared from the region, while
they persist in the eastern forests to this
day.
The third type of forest which occurs in the
eastern half of North America is the “rain
forest” of Florida and parts of the adjacent
states. Here the rainfall is abundant all the
year round, with a summer maximum, and
the temperature is high. There is thus no
need to economise water, and where the soil
permits there is a luxuriant type of forest,
which recalls that of the tropics, although it
is poorer. Where soil conditions are unfavourable
we have pine woods, conifers
throughout the eastern United States always
taking advantage of conditions relatively
unfavourable to the broad-leafed trees.
Thus if we follow the eastern seaboard
of the United States from Labrador to Florida
we pass through the following floral regions:—(1)
Coniferous forest, with relatively few
species, (2) mixed coniferous and deciduous
forest with chiefly the harder types of
deciduous trees, (3) predominantly deciduous
forest with many of the larger-leafed and
more delicate forms, and finally (4) forest of
the sub-tropical rainy type, intermixed with
coniferous woods on the barren sandy soil
and in the swamps.
The western coast shows more uniformity,
the western type of coniferous forest stretching
from Alaska to California, though it is
richer, and more luxuriant in the warmer
regions when moisture is still obtainable.
As the moisture diminishes the forest dies
away and desert or semi-desert conditions
supervene.