I have said enough to show that Mr.
Darwin claimed I to have been the originator of the theory of
descent with modification as distinctly as any writer usually
claims any theory; but it will probably save the reader trouble
in the end if I bring together a good many, though not, probably,
all (for I much disliked the task, and discharged it
perfunctorily), of the passages in the “Origin of
Species” in which the theory of descent with modification
in its widest sense is claimed expressly or by implication.
I shall quote from the original edition, which, it should be
remembered, consisted of the very unusually large number of four
thousand copies, and from which no important deviation was made
either by addition or otherwise until a second edition of two
thousand further copies had been sold; the “Historical
Sketch,” &c., being first given with the third
edition. The italics, which I have employed so as to catch
the reader’s eye, are mine, not Mr. Darwin’s.
Mr. Darwin writes:—
“Although much remains obscure, and will long remain
obscure, I can entertain no doubt, after the most
deliberate study and dispassionate judgment of which I am
capable, that the view which most naturalists
entertain, and which I formerly entertained—namely
that each species has been independently created—is
erroneous. I am fully convinced that species are not
immutable, but that those belonging to what are called the same
genera are lineal descendants of some other and generally extinct
species, in the same manner as the acknowledged varieties of any
one species are the descendants of that species.
Furthermore, I am convinced that natural selection” (or the
preservation of fortunate races) “has been the main but not
exclusive means of modification” (p. 6).
It is not here expressly stated that the theory of the
mutability of species is Mr. Darwin’s own; this,
nevertheless, is the inference which the great majority of his
readers were likely to draw, and did draw, from Mr.
Darwin’s words.
Again:—
“It is not that all large genera are now varying much,
and are thus increasing in the number of their species, or that
no small genera are now multiplying and increasing; for if this
had been so it would have been fatal to my theory;
inasmuch as geology,” &c. (p. 56).
The words “my theory” stand in all the
editions. Again:—
“This relation has a clear meaning on my view of
the subject; I look upon all the species of any genus as having
as certainly descended from the same progenitor, as have the two
sexes of any one of the species” (p. 157).
“My view” here, especially in the absence of
reference to any other writer as having held the same opinion,
implies as its most natural interpretation that descent pure and
simple is Mr. Darwin’s view. Substitute “the
theory of descent” for “my view,” and we do not
feel that we are misinterpreting the author’s
meaning. The words “my view” remain in all
editions.
Again:—
“Long before having arrived at this part of my work, a
crowd of difficulties will have occurred to the reader.
Some of them are so grave that to this day I can never reflect on
them without being staggered; but to the best of my belief the
greater number are only apparent, and those that are real are
not, I think, fatal to my theory.
“These difficulties and objections may be classed under
the following heads:—Firstly, if species have descended
from other species by insensibly fine gradations, why do we not
everywhere see?” &c. (p. 171).
We infer from this that “my theory” is the theory
“that species have descended from other species by
insensibly fine gradations”—that is to say, that it
is the theory of descent with modification; for the theory that
is being objected to is obviously the theory of descent in
toto, and not a mere detail in connection with that
theory.
The words “my theory” were altered in 1872, with
the sixth edition of the “Origin of species,” into
“the theory;” but I am chiefly concerned with the
first edition of the work, my object being to show that Mr.
Darwin was led into his false position as regards natural
selection by a desire to claim the theory of descent with
modification; if he claimed it in the first edition, this is
enough to give colour to the view which I take; but it must be
remembered that descent with modification remained, by the
passage just quoted “my theory,” for thirteen years,
and even when in 1869 and 1872, for a reason that I can only
guess at, “my theory” became generally “the
theory,” this did not make it become any one else’s
theory. It is hard to say whose or what it became, if the
words are to be construed technically; practically, however, with
all ingenuous readers, “the theory” remained as much
Mr. Darwin’s theory as though the words “my
theory” had been retained, and Mr. Darwin cannot be
supposed so simple-minded as not to have known this would be the
case. Moreover, it appears, from the next page but one to
the one last quoted, that Mr. Darwin claimed the theory of
descent with modification generally, even to the last, for we
there read, “By my theory these allied species have
descended from a common parent,” and the “my”
has been allowed, for some reason not quite obvious, to survive
the general massacre of Mr. Darwin’s
“my’s” which occurred in 1869 and 1872.
Again:—
“He who believes that each being has been created as we
now see it, must occasionally have felt surprise when he has
met,” &c. (p. 185).
Here the argument evidently lies between descent and
independent acts of creation. This appears from the
paragraph immediately following, which begins, “He who
believes in separate and innumerable acts of creation,”
&c. We therefore understand descent to be the theory so
frequently spoken of by Mr. Darwin as “my.”
Again:—
“He who will go thus far, if he find on finishing this
treatise that large bodies of facts, otherwise inexplicable, can
be explained by the theory of descent, ought not to
hesitate to go farther, and to admit that a structure even as
perfect as an eagle’s eye might be formed by natural
selection, although in this case he does not know any of the
transitional grades” (p. 188).
The natural inference from this is that descent and natural
selection are one and the same thing.
Again:—
“If it could be demonstrated that any complex organ
existed which could not possibly have been formed by numerous,
successive, slight modifications, my theory would
absolutely break down. But I can find out no such
case. No doubt many organs exist of which we do not know
the transitional grades, more especially if we look to
much-isolated species, round which, according to my
theory, there has been much extinction” (p.
189).
This makes “my theory” to be “the theory
that complex organs have arisen by numerous, successive, slight
modifications;” that is to say, to be the theory of descent
with modification. The first of the two “my
theory’s” in the passage last quoted has been allowed
to stand. The second became “the theory” in
1872. It is obvious, therefore, that “the
theory” means “my theory;” it is not so obvious
why the change should have been made at all, nor why the one
“my theory” should have been taken and the other
left, but I will return to this question.
Again, Mr. Darwin writes:—
“Although we must be extremely cautious in concluding
that any organ could not possibly have been produced by small
successive transitional gradations, yet, undoubtedly grave cases
of difficulty occur, some of which will be discussed in my future
work” (p. 192).
This, as usual, implies descent with modification to be the
theory that Mr. Darwin is trying to make good.
Again:—
“I have been astonished how rarely an organ can be named
towards which no transitional variety is known to lead . . . Why,
on the theory of creation, should this be so? Why
should not nature have taken a leap from structure to
structure? On the theory of natural selection we can
clearly understand why she should not; for natural selection can
act only by taking advantage of slight successive variations; she
can never take a leap, but must advance by the slowest and
shortest steps” (p. 194).
Here “the theory of natural selection” is opposed
to “the theory of creation;” we took it, therefore,
to be another way of saying “the theory of descent with
modification.”
Again:—
“We have in this chapter discussed some of the
difficulties and objections which may be urged against my
theory. Many of them are very grave, but I think that
in the discussion light has been thrown on several facts which,
on the theory of independent acts of creation, are utterly
obscure” (p. 203).
Here we have, on the one hand, “my theory,” on the
other, “independent acts of creation.” The
natural antithesis to independent acts of creation is descent,
and we assumed with reason that Mr. Darwin was claiming this when
he spoke of “my theory.” “My
theory” became “the theory” in 1869.
Again:—
“On the theory of natural selection we can clearly
understand the full meaning of that old canon in natural history,
‘Natura non facit saltum.’ This canon,
if we look only to the present inhabitants of the world is not
strictly correct, but if we include all those of past times, it
must by my theory be strictly true” (p. 206).
Here the natural interpretation of “by my theory”
is “by the theory of descent with modification;” the
words “on the theory of natural selection,” with
which the sentence opens, lead us to suppose that Mr. Darwin
regarded natural selection and descent as convertible
terms. “My theory” was altered to “this
theory” in 1872. Six lines lower down we read,
“On my theory unity of type is explained by unity of
descent.” The “my” here has been allowed
to stand.
Again:—
“Again, as in the case of corporeal structure, and
conformably with my theory, the instinct of each species
is good for itself, but has never,” &c. (p. 210).
Who was to see that “my theory” did not include
descent with modification? The “my” here has
been allowed to stand.
Again:—
“The fact that instincts . . . are liable to make
mistakes;—that no instinct has been produced for the
exclusive good of other animals, but that each animal takes
advantage of the instincts of others;—that the canon of
natural history, ‘Natura non facit saltum,’ is
applicable to instincts as well as to corporeal structure, and is
plainly explicable on the foregoing views, but is otherwise
inexplicable,—all tend to corroborate the theory of
natural selection” (p. 243).
We feel that it is the theory of evolution, or descent with
modification, that is here corroborated, and that it is this
which Mr. Darwin is mainly trying to establish; the sentence
should have ended “all tend to corroborate the theory of
descent with modification;” the substitution of
“natural selection” for descent tends to make us
think that these conceptions are identical. That they are
so regarded, or at any rate that it is the theory of descent in
full which Mr. Darwin has in his mind, appears from the
immediately succeeding paragraph, which begins “This
theory,” and continues six lines lower, “For
instance, we can understand, on the principle of
inheritance, how it is that,” &c.
Again:—
“In the first place, it should always be borne in mind
what sort of intermediate forms must, on my theory,
formerly have existed” (p. 280).
“My theory” became “the theory” in
1869. No reader who read in good faith could doubt that the
theory of descent with modification was being here intended.
“It is just possible by my theory, that one of
two living forms might have descended from the other; for
instance, a horse from a tapir; but in this case direct
intermediate links will have existed between them” (p.
281).
“My theory” became “the theory” in
1869.
Again:—
“By the theory of natural selection all living
species have been connected with the parent species of each
genus,” &c. We took this to mean, “By the
theory of descent with modification all living species,”
&c. (p. 281).
Again:—
“Some experienced conchologists are now sinking many of
the very fine species of D’Orbigny and others into the rank
of varieties; and on this view we do find the kind of evidence of
change which on my theory we ought to find” (p.
297).
“My theory” became “the theory” in
1869.
In the fourth edition (1866), in a passage which is not in
either of the two first editions, we read (p. 359), “So
that here again we have undoubted evidence of change in the
direction required by my theory.” “My
theory” became “the theory” in 1869; the theory
of descent with modification is unquestionably intended.
Again:—
“Geological research has done scarcely anything in
breaking down the distinction between species, by connecting them
together by numerous, fine, intermediate varieties; and this not
having been effected, is probably the gravest and most obvious of
all the many objections which may be urged against my
views” (p. 299).
We naturally took “my views” to mean descent with
modification. The “my” has been allowed to
stand.
Again:—
“If, then, there be some degree of truth in these
remarks, we have no right to expect to find in our geological
formations an infinite number of those transitional forms which
on my theory assuredly have connected all the past and
present species of the same group in one long and branching chain
of life . . . But I do not pretend that I should ever have
suspected how poor was the record in the best preserved
geological sections, had not the absence of innumerable
transitional links between the species which lived at the
commencement and at the close of each formation pressed so hardly
on my theory” (pp. 301, 302).
Substitute “descent with modification” for
“my theory” and the meaning does not suffer.
The first of the two “my theories” in the passage
last quoted was altered in 1869 into “our theory;”
the second has been allowed to stand.
Again:—
“The abrupt manner in which whole groups of species
suddenly appear in some formations, has been urged by several
palæontologists . . . as a fatal objection to the belief
in the transmutation of species. If numerous species,
belonging to the same genera or families, have really started
into life all at once, the fact would be fatal to the theory
of descent with slow modification through natural
selection” (p. 302).
Here “the belief in the transmutation of species,”
or descent with modification, is treated as synonymous with
“the theory of descent with slow modification through
natural selection;” but it has nowhere been explained that
there are two widely different “theories of descent with
slow modification through natural selection,” the one of
which may be true enough for all practical purposes, while the
other is seen to be absurd as soon as it is examined
closely. The theory of descent with modification is not
properly convertible with either of these two views, for descent
with modification deals with the question whether species are
transmutable or no, and dispute as to the respective merits of
the two natural selections deals with the question how it comes
to be transmuted; nevertheless, the words “the theory of
descent with slow modification through the ordinary course of
things” (which is what “descent with modification
through natural selection” comes to) may be considered as
expressing the facts with practical accuracy, if the ordinary
course of nature is supposed to be that modification is mainly
consequent on the discharge of some correlated function, and that
modification, if favourable, will tend to accumulate so long as
the given function continues important to the wellbeing of the
organism; the words, however, have no correspondence with reality
if they are supposed to imply that variations which are mainly
matters of pure chance and unconnected in any way with function
will accumulate and result in specific difference, no matter how
much each one of them may be preserved in the generation in which
it appears. In the one case, therefore, the expression
natural selection may be loosely used as a synonym for descent
with modification, and in the other it may not.
Unfortunately with Mr. Charles Darwin the variations are mainly
accidental. The words “through natural
selection,” therefore, in the passage last quoted carry no
weight, for it is the wrong natural selection that is, or ought
to be, intended; practically, however, they derived a weight from
Mr. Darwin’s name to which they had no title of their own,
and we understood that “the theory of descent with slow
modification” through the kind of natural selection
ostensibly intended by Mr. Darwin was a quasi-synonymous
expression for the transmutation of species. We
understood—so far as we understood anything beyond that we
were to believe in descent with modification—that natural
selection was Mr. Darwin’s theory; we therefore concluded,
since Mr. Darwin seemed to say so, that the theory of the
transmutation of species generally was so also. At any rate
we felt as regards the passage last quoted that the theory of
descent with modification was the point of attack and defence,
and we supposed it to be the theory so often referred to by Mr.
Darwin as “my.”
Again:—
“Some of the most ancient Silurian animals, as the
Nautilus, Lingula, &c., do not differ much from the living
species; and it cannot on my theory be supposed that these
old species were the progenitors,” &c. (p. 306) . . .
“Consequently if my theory be true, it is
indisputable,” &c. (p. 307).
Here the two “my theories” have been altered, the
first into “our theory,” and the second into
“the theory,” both in 1869; but, as usual, the thing
that remains with the reader is the theory of descent, and it
remains morally and practically as much claimed when called
“the theory”—as during the many years
throughout which the more open “my” distinctly
claimed it.
Again:—
“All the most eminent palæontologists, namely,
Cuvier, Owen, Agassiz, Barrande, E. Forbes, &c., and all our
greatest geologists, as Lyell, Murchison, Sedgwick, &c., have
unanimously, often vehemently, maintained the immutability of
species. . . . I feel how rash it is to differ from these
great authorities . . . Those who think the natural geological
record in any degree perfect, and who do not attach much weight
to the facts and arguments of other kinds brought forward in this
volume, will undoubtedly at once reject my theory”
(p. 310).
What is “my theory” here, if not that of the
mutability of species, or the theory of descent with
modification? “My theory” became “the
theory” in 1869.
Again:—
“Let us now see whether the several facts and rules
relating to the geological succession of organic beings, better
accord with the common view of the immutability of species, or
with that of their slow and gradual modification,
through descent and natural selection” (p. 312).
The words “natural selection” are indeed here, but
they might as well be omitted for all the effect they
produce. The argument is felt to be about the two opposed
theories of descent, and independent creative efforts.
Again:—
“These several facts accord well with my
theory” (p. 314). That “my theory” is
the theory of descent is the conclusion most naturally drawn from
the context. “My theory” became “our
theory” in 1869.
Again:—
“This gradual increase in the number of the species of a
group is strictly conformable with my theory; for the
process of modification and the production of a number of allied
forms must be slow and gradual, . . . like the branching of a
great tree from a single stem, till the group becomes
large” (p. 314).
“My theory” became “the theory” in
1869. We took “my theory” to be the theory of
descent; that Mr. Darwin treats this as synonymous with the
theory of natural selection appears from the next paragraph, on
the third line of which we read, “On the theory of
natural selection the extinction of old forms,”
&c.
Again:—
“The theory of natural selection is grounded on
the belief that each new variety and ultimately each new species,
is produced and maintained by having some advantage over those
with which it comes into competition; and the consequent
extinction of less favoured forms almost inevitably
follows” (p. 320). Sense and consistency cannot be
made of this passage. Substitute “The theory of the
preservation of favoured races in the struggle for life”
for “The theory of natural selection” (to do this is
only taking Mr. Darwin’s own synonym for natural selection)
and see what the passage comes to. “The preservation
of favoured races” is not a theory, it is a commonly
observed fact; it is not “grounded on the belief that each
new variety,” &c., it is one of the ultimate and most
elementary principles in the world of life. When we try to
take the passage seriously and think it out, we soon give it up,
and pass on, substituting “the theory of descent” for
“the theory of natural selection,” and concluding
that in some way these two things must be identical.
Again:—
“The manner in which single species and whole groups of
species become extinct accords well with the theory of natural
selection” (p. 322).
Again:—
“This great fact of the parallel succession of the forms
of life throughout the world, is explicable on the theory of
natural selection” (p. 325).
Again:—
“Let us now look to the mutual affinities of extinct and
living species. They all fall into one grand natural
system; and this is at once explained on the principle of
descent” (p. 329).
Putting the three preceding passages together, we naturally
inferred that “the theory of natural selection” and
“the principle of descent” were the same
things. We knew Mr. Darwin claimed the first, and therefore
unhesitatingly gave him the second at the same time.
Again:—
“Let us see how far these several facts and inferences
accord with the theory of descent with modification”
(p. 331)
Again:—
“Thus, on the theory of descent with
modification, the main facts with regard to the mutual
affinities of the extinct forms of life to each other and to
living forms, seem to me explained in a satisfactory
manner. And they are wholly inexplicable on any other
view” (p. 333).
The words “seem to me” involve a claim in the
absence of so much as a hint in any part of the book concerning
indebtedness to earlier writers.
Again:—
“On the theory of descent, the full meaning of
the fossil remains,” &c. (p. 336).
In the following paragraph we read:—
“But in one particular sense the more recent forms must,
on my theory, be higher than the more ancient.”
Again:—
“Agassiz insists that ancient animals resemble to a
certain extent the embryos of recent animals of the same classes;
or that the geological succession of extinct forms is in some
degree parallel to the embryological development of recent forms.
. . . This doctrine of Agassiz accords well with the theory of
natural selection” (p. 338).
“The theory of natural selection” became
“our theory” in 1869. The opinion of Agassiz
accords excellently with the theory of descent with modification,
but it is not easy to see how it bears upon the fact that lucky
races are preserved in the struggle for life—which,
according to Mr. Darwin’s title-page, is what is meant by
natural selection.
Again:—
“On the theory of descent with modification, the
great law of the long-enduring but not immutable succession of
the same types within the same areas, is at once explained”
(p. 340).
Again:—
“It must not be forgotten that, on my theory, all
the species of the same genus have descended from some one
species” (p. 341).
“My theory” became “our theory” in
1869.
Again:—
“He who rejects these views on the nature of the
geological record, will rightly reject my whole
theory” (p. 342).
“My” became “our” in 1869.
Again:—
“Passing from these difficulties, the other great
leading facts in palæontology agree admirably with the
theory of descent with modification through variation and natural
selection” (p. 343).
Again:—
The succession of the same types of structure within the same
areas during the later geological periods ceases to be
mysterious, and is simply explained by inheritance (p.
345).
I suppose inheritance was not when Mr. Darwin wrote considered
mysterious. The last few words have been altered to
“and is intelligible on the principle of
inheritance.” It seems as though Mr. Darwin did not
like saying that inheritance was not mysterious, but had no
objection to implying that it was intelligible.
The next paragraph begins—“If, then, the
geological record be as imperfect as I believe it to be, . . .
the main objections to the theory of natural selection are
greatly diminished or disappear. On the other hand, all the
chief laws of palæontology plainly proclaim, as it seems
to me, that species have been produced by ordinary
generation.”
Here again the claim to the theory of descent with
modification is unmistakable; it cannot, moreover, but occur to
us that if species “have been produced by ordinary
generation,” then ordinary generation has as good a claim
to be the main means of originating species as natural selection
has. It is hardly necessary to point out that ordinary
generation involves descent with modification, for all known
offspring differ from their parents, so far, at any rate, as that
practised judges can generally tell them apart.
Again:—
“We see in these facts some deep organic bond,
prevailing throughout space and time, over the same areas of land
and water, and independent of their physical condition. The
naturalist must feel little curiosity who is not led to inquire
what this bond is.
“This bond, on my theory, is simply
inheritance, that cause which alone,” &c. (p.
350).
This passage was altered in 1869 to “The bond is simply
inheritance.” The paragraph concludes, “On
this principle of inheritance with modification, we can
understand how it is that sections of genera . . . are confined
to the same areas,” &c.
Again:—
“He who rejects it rejects the vera causa of
ordinary generation,” &c. (p. 352).
We naturally ask, Why call natural selection the “main
means of modification,” if “ordinary
generation” is a vera causa?
Again:—
“In discussing this subject, we shall be enabled at the
same time to consider a point equally important for us, namely,
whether the several distinct species of a genus, which on my
theory have all descended from a common ancestor, can have
migrated (undergoing modification during some part of their
migration) from the area inhabited by their progenitor” (p.
354).
The words “on my theory” became “on our
theory” in 1869.
Again:—
“With those organic beings which never intercross (if
such exist) the species, on my theory, must have
descended from a succession of improved varieties,”
&c. (p. 355).
The words “on my theory” were cut out in 1869.
Again:—
“A slow southern migration of a marine fauna will
account, on the theory of modification, for many closely
allied forms,” &c. (p. 372).
Again:—
“But the existence of several quite distinct species,
belonging to genera exclusively confined to the southern
hemisphere, is, on my theory of descent with modification,
a far more remarkable case of difficulty” (p. 381).
“My” became “the” in 1866 with the
fourth edition. This was the most categorical claim to the
theory of descent with modification in the “Origin of
Species.” The “my” here is the only one
that was taken out before 1869. I suppose Mr. Darwin
thought that with the removal of this “my” he had
ceased to claim the theory of descent with modification.
Nothing, however, could be gained by calling the reader’s
attention to what had been done, so nothing was said about
it.
Again:—
“Some species of fresh-water shells have a very wide
range, and allied species, which, on my
theory, are descended from a single source, prevail
throughout the world” (p. 385).
“My theory” became “our theory” in
1869.
Again:—
“In the following remarks I shall not confine myself to
the mere question of dispersal, but shall consider some other
facts which bear upon the truth of the two theories of
independent creation and of descent with modification”
(p. 389). What can be plainer than that the theory which
Mr. Darwin espouses, and has so frequently called
“my,” is descent with modification?
Again:—
“But as these animals and their spawn are known to be
immediately killed by sea-water, on my view, we can see
that there would be great difficulty in their transportal across
the sea, and therefore why they do not exist on any oceanic
island. But why, on the theory of creation, they
should not have been created there, it would be very difficult to
explain” (p. 393).
“On my view” was cut out in 1869.
On the following page we read—“On my view this
question can easily be answered.” “On my
view” is retained in the latest edition.
Again:—
“Yet there must be, on my view, some unknown but
highly efficient means for their transportation” (p.
397).
“On my view” became “according to our
view” in 1869.
Again:—
“I believe this grand fact can receive no sort of
explanation on the ordinary view of independent creation;
whereas, on the view here maintained, it is obvious that
the Galapagos Islands would be likely to receive colonists . . .
from America, and the Cape de Verde Islands from Africa; and that
such colonists would be liable to modification; the principle of
inheritance still betraying their original birth-place” (p.
399).
Again:—
“With respect to the distinct species of the same genus
which, on my theory, must have spread from one parent
source, if we make the same allowances as before,”
&c.
“On my theory” became “on our theory”
in 1869.
Again:—
“On my theory these several relations throughout
time and space are intelligible; . . . the forms within each
class have been connected by the same bond of ordinary
generation; . . . in both cases the laws of variation have been
the same, and modifications have been accumulated by the same
power of natural selection” (p. 410).
“On my theory” became “according to our
theory” in 1869, and natural selection is no longer a
power, but has become a means.
Again:—
“I believe that something more is included, and
that propinquity of descent—the only known cause of the
similarity of organic beings—is the bond, hidden as it is
by various degrees of modification, which is partially revealed
to us by our classification” (p. 418).
Again:—
“Thus, on the view which I hold, the
natural system is genealogical in its arrangement, like a
pedigree” (p. 422).
“On the view which I hold” was cut out in
1872.
Again:—
“We may feel almost sure, on the theory of
descent, that these characters have been inherited from a
common ancestor” (p. 426).
Again:—
“On my view of characters being of real importance
for classification only in so far as they reveal descent, we
can clearly understand,” &c. (p. 427).
“On my view” became “on the view” in
1872.
Again:—
“The more aberrant any form is, the greater must be the
number of connecting forms which, on my theory, have been
exterminated and utterly lost” (p. 429).
The words “on my theory” were excised in 1869.
Again:—
“Finally, we have seen that natural selection
. . . explains that great and universal feature in the
affinities of all organic beings, namely, their subordination in
group under group. We use the element of descent in
classing the individuals of both sexes, &c.; . . . we use
descent in classing acknowledged varieties; . . . and I
believe this element of descent is the hidden bond of connection
which naturalists have sought under the term of the natural
system” (p. 433).
Lamarck was of much the same opinion, as I showed in
“Evolution Old and New.” He
wrote:—“An arrangement should be considered
systematic, or arbitrary, when it does not conform to the
genealogical order taken by nature in the development of the
things arranged, and when, by consequence, it is not founded on
well-considered analogies. There is a natural order in
every department of nature; it is the order in which its several
component items have been successively developed.” [195a] The point, however, which
should more particularly engage our attention is that Mr. Darwin
in the passage last quoted uses “natural selection”
and “descent” as though they were convertible
terms.
Again:—
“Nothing can be more hopeless than to attempt to explain
this similarity of pattern in members of the same class by
utility or the doctrine of final causes . . . On the
ordinary view of the independent creation of each being, we
can only say that so it is . . . The explanation is manifest
on the theory of the natural selection of successive slight
modifications,” &c. (p. 435).
This now stands—“The explanation is to a large
extent simple, on the theory of the selection of successive,
slight modifications.” I do not like “a large
extent” of simplicity; but, waiving this, the point at
issue is not whether the ordinary course of things ensures a
quasi-selection of the types that are best adapted to their
surroundings, with accumulation of modification in various
directions, and hence wide eventual difference between species
descended from common progenitors—no evolutionist since
1750 has doubted this—but whether a general principle
underlies the modifications from among which the quasi-selection
is made, or whether they are destitute of such principle and
referable, as far as we are concerned, to chance only.
Waiving this again, we note that the theories of independent
creation and of natural selection are contrasted, as though they
were the only two alternatives; knowing the two alternatives to
be independent creation and descent with modification, we
naturally took natural selection to mean descent with
modification.
Again:—
“On the theory of natural selection we can
satisfactorily answer these questions” (p. 437).
“Satisfactorily” now stands “to a certain
extent.”
Again:—
“On my view these terms may be used
literally” (pp. 438, 439).
“On my view” became “according to the views
here maintained such language may be,” &c., in
1869.
Again:—
“I believe all these facts can be explained as follows,
on the view of descent with modification” (p.
443).
This sentence now ends at “follows.”
Again:—
“Let us take a genus of birds, descended, on
my theory, from some one parent species, and of which
the several new species have become modified through natural
selection in accordance with their divers habits” (p.
446).
The words “on my theory” were cut out in 1869, and
the passage now stands, “Let us take a group of birds,
descended from some ancient form and modified through natural
selection for different habits.”
Again:—
“On my view of descent with modification, the
origin of rudimentary organs is simple” (p. 454).
“On my view” became “on the
view” in 1869.
Again:—
“On the view of descent with modification,”
&c. (p. 455).
Again:—
“On this same view of descent with modification
all the great facts of morphology become intelligible” (p.
456).
Again:—
“That many and grave objections may be advanced against
the theory of descent with modification through natural
selection, I do not deny” (p. 459).
This now stands, “That many and serious objections may
be advanced against the theory of descent with modification
through variation and natural selection, I do not
deny.”
Again:—
“There are, it must be admitted, cases of special
difficulty on the theory of natural selection” (p.
460).
“On” has become “opposed to;” it is
not easy to see why this alteration was made, unless because
“opposed to” is longer.
Again:—
“Turning to geographical distribution, the difficulties
encountered on the theory of descent with modification are
grave enough.”
“Grave” has become “serious,” but
there is no other change (p. 461).
Again:—
“As on the theory of natural selection an
interminable number of intermediate forms must have
existed,” &c.
“On” has become “according
to”—which is certainly longer, but does not appear to
possess any other advantage over “on.” It is
not easy to understand why Mr. Darwin should have strained at
such a gnat as “on,” though feeling no discomfort in
such an expression as “an interminable number.”
Again:—
“This is the most forcible of the many objections which
may be urged against my theory . . . For certainly, on
my theory,” &c. (p. 463).
The “my” in each case became “the” in
1869.
Again:—
“Such is the sum of the several chief objections and
difficulties which may be justly urged against my
theory” (p. 465).
“My” became “the” in 1869.
Again:—
“Grave as these several difficulties are, in my
judgment they do not overthrow the theory of descent with
modifications” (p. 466).
This now stands, “Serious as these several objections
are, in my judgment they are by no means sufficient to overthrow
the theory of descent with subsequent modification;”
which, again, is longer, and shows at what little, little gnats
Mr. Darwin could strain, but is no material amendment on the
original passage.
Again:—
“The theory of natural selection, even if we
looked no further than this, seems to me to be in itself
probable” (p. 469).
This now stands, “The theory of natural selection, even
if we look no further than this, seems to be in the highest
degree probable.” It is not only probable, but
was very sufficiently proved long before Mr. Darwin was born,
only it must be the right natural selection and not Mr. Charles
Darwin’s.
Again:—
“It is inexplicable, on the theory of creation,
why a part developed, &c., . . . but, on my
view, this part has undergone,” &c. (p. 474).
“On my view” became “on our view” in
1869.
Again:—
“Glancing at instincts, marvellous as some are, they
offer no greater difficulty than does corporeal structure on
the theory of the natural selection of successive,
slight, but profitable modifications” (p.
474).
Again:—
“On the view of all the species of the same genus
having descended from a common parent, and having inherited
much in common, we can understand how it is,” &c. (p.
474).
Again:—
“If we admit that the geological record is imperfect in
an extreme degree, then such facts as the record gives, support
the theory of descent with modification.
“ . . . The extinction of species . . . almost
inevitably follows on the principle of natural
selection” (p. 475).
The word “almost” has got a great deal to answer
for.
Again:—
“We can understand, on the theory of descent with
modification, most of the great leading facts in
Distribution” (p. 476).
Again:—
“The existence of closely allied or representative
species in any two areas, implies, on the theory of descent
with modification, that the same parents formerly inhabited
both areas . . . It must be admitted that these facts receive no
explanation on the theory of creation . . . The fact . . .
is intelligible on the theory of natural selection, with
its contingencies of extinction and divergence of
character” (p. 478).
Again:—
“Innumerable other such facts at once explain themselves
on the theory of descent with slow and slight successive
modifications” (p. 479).
“Any one whose disposition leads him to attach more
weight to unexplained difficulties than to the explanation of a
certain number of facts, will certainly reject my
theory” (p. 482).
“My theory” became “the theory” in
1869.
From this point to the end of the book the claim is so
ubiquitous, either expressly or by implication, that it is
difficult to know what not to quote. I must, however,
content myself with only a few more extracts. Mr. Darwin
says:—
“It may be asked how far I extend the doctrine of the
modification of species” (p. 482).
Again:—
“Analogy would lead me one step further, namely, to the
belief that all animals and plants have descended from some one
prototype . . . Therefore I should infer from analogy that
probably all the organic beings which have ever lived on this
earth have descended from some one primordial form, into which
life was first breathed.”
From an amœba—Adam, in fact, though not in
name. This last sentence is now completely altered, as well
it might be.
Again:—
“When the views entertained in this volume on the
origin of species, or when analogous views are generally
admitted, we can dimly foresee that there will be a
considerable revolution in natural history” (p. 434).
Possibly. This now stands, “When the views
advanced by me in this volume, and by Mr. Wallace, or when
analogous views on the origin of species are generally admitted,
we can dimly foresee,” &c. When the “Origin
of Species” came out we knew nothing of any analogous
views, and Mr. Darwin’s words passed unnoticed. I do
not say that he knew they would, but he certainly ought to have
known.
Again:—
“A grand and almost untrodden field of inquiry will
be opened, on the causes and laws of variation, on
correlation of growth, on the effects of use and disuse, on the
direct action of external conditions, and so forth” (p.
486).
Buffon and Lamarck had trodden this field to some purpose, but
not a hint to this effect is vouchsafed to us.
Again;—
“When I view all beings not as special creations,
but as the lineal descendants of some few beings which lived
long before the first bed of the Silurian system was
deposited, they seem to me to become ennobled . . . We can so far
take a prophetic glance into futurity as to foretell that it will
be the common and widely spread species, belonging to the larger
and dominant groups, which will ultimately prevail and procreate
new and dominant species.”
There is no alteration in this except that
“Silurian” has become “Cambrian.”
The idyllic paragraph with which Mr. Darwin concludes his book
contains no more special claim to the theory of descent en
bloc than many another which I have allowed to pass
unnoticed; it has been, moreover, dealt with in an earlier
chapter (Chapter XII.)